Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea (Ukraine)
Twenty species of the Pselaphinae (Coleoptera, Staphylinidae) are recorded for the first time from Crimean Peninsula. Bryaxis moczarskii (Winkler, 1911) is redescribed and illustrated for the first time after its original description. Brachygluta guillemardi (Saulcy, 1876), Bryaxis longulus (Kiesenw...
Збережено в:
| Дата: | 2013 |
|---|---|
| Автор: | |
| Формат: | Стаття |
| Мова: | English |
| Опубліковано: |
Інститут зоології ім. І.І. Шмальгаузена НАН України
2013
|
| Назва видання: | Вестник зоологии |
| Теми: | |
| Онлайн доступ: | http://dspace.nbuv.gov.ua/handle/123456789/109791 |
| Теги: |
Додати тег
Немає тегів, Будьте першим, хто поставить тег для цього запису!
|
| Назва журналу: | Digital Library of Periodicals of National Academy of Sciences of Ukraine |
| Цитувати: | Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea (Ukraine) / R.E. Krivosheyev // Вестник зоологии. — 2013. — Т. 47, № 4. — С. 299–307. — Бібліогр.: 15 назв. — англ. |
Репозитарії
Digital Library of Periodicals of National Academy of Sciences of Ukraine| id |
irk-123456789-109791 |
|---|---|
| record_format |
dspace |
| spelling |
irk-123456789-1097912016-12-15T03:02:24Z Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea (Ukraine) Krivosheyev, R.E. Фауна и систематика Twenty species of the Pselaphinae (Coleoptera, Staphylinidae) are recorded for the first time from Crimean Peninsula. Bryaxis moczarskii (Winkler, 1911) is redescribed and illustrated for the first time after its original description. Brachygluta guillemardi (Saulcy, 1876), Bryaxis longulus (Kiesenwetter, 1849) and Claviger colchicus Motschulsky, 1837 are recorded for the first time from Ukraine. In total, 34 species of ten genera and one unrecognizable nominal species are recorded from Crimea. Впервые для Крымского полуострова указаны 20 видов ощупников. Приведено иллюстрированное переописание Bryaxis moczarskii (Winkler, 1911). Brachygluta guillemardi (Saulcy, 1876), Bryaxis longulus (Kiesenwetter, 1849) и Claviger colchicus Motschulsky, 1837 отмечены впервые в фауне Украины. В целом из Крыма приведены 34 вида из 10 родов и один не подлежащий идентификации номинальный вид. 2013 Article Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea (Ukraine) / R.E. Krivosheyev // Вестник зоологии. — 2013. — Т. 47, № 4. — С. 299–307. — Бібліогр.: 15 назв. — англ. 0084-5604 DOI 10.2478/vzoo-2013-0031 http://dspace.nbuv.gov.ua/handle/123456789/109791 595.763.34(282.247.34) en Вестник зоологии Інститут зоології ім. І.І. Шмальгаузена НАН України |
| institution |
Digital Library of Periodicals of National Academy of Sciences of Ukraine |
| collection |
DSpace DC |
| language |
English |
| topic |
Фауна и систематика Фауна и систематика |
| spellingShingle |
Фауна и систематика Фауна и систематика Krivosheyev, R.E. Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea (Ukraine) Вестник зоологии |
| description |
Twenty species of the Pselaphinae (Coleoptera, Staphylinidae) are recorded for the first time from Crimean Peninsula. Bryaxis moczarskii (Winkler, 1911) is redescribed and illustrated for the first time after its original description. Brachygluta guillemardi (Saulcy, 1876), Bryaxis longulus (Kiesenwetter, 1849) and Claviger colchicus Motschulsky, 1837 are recorded for the first time from Ukraine. In total, 34 species of ten genera and one unrecognizable nominal species are recorded from Crimea. |
| format |
Article |
| author |
Krivosheyev, R.E. |
| author_facet |
Krivosheyev, R.E. |
| author_sort |
Krivosheyev, R.E. |
| title |
Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea (Ukraine) |
| title_short |
Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea (Ukraine) |
| title_full |
Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea (Ukraine) |
| title_fullStr |
Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea (Ukraine) |
| title_full_unstemmed |
Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea (Ukraine) |
| title_sort |
review of the short-winged mold beetles (coleoptera, staphylinidae, pselaphinae) of the crimea (ukraine) |
| publisher |
Інститут зоології ім. І.І. Шмальгаузена НАН України |
| publishDate |
2013 |
| topic_facet |
Фауна и систематика |
| url |
http://dspace.nbuv.gov.ua/handle/123456789/109791 |
| citation_txt |
Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea (Ukraine) / R.E. Krivosheyev // Вестник зоологии. — 2013. — Т. 47, № 4. — С. 299–307. — Бібліогр.: 15 назв. — англ. |
| series |
Вестник зоологии |
| work_keys_str_mv |
AT krivosheyevre reviewoftheshortwingedmoldbeetlescoleopterastaphylinidaepselaphinaeofthecrimeaukraine |
| first_indexed |
2025-07-07T23:39:52Z |
| last_indexed |
2025-07-07T23:39:52Z |
| _version_ |
1837033424454418432 |
| fulltext |
UDC 595.763.34(282.247.34)
REVIEW OF THE SHORT-WINGED MOLD BEETLES
(COLEOPTERA, STAPHYLINIDAE, PSELAPHINAE)
OF THE CRIMEA (UKRAINE)
R. E. Krivosheyev
Schmalhausen Institute of Zoology NAS of Ukraine,
vul. B. Khmelnytskogo, 15, Kyiv, 01601 Ukraine
E-mail: accipitergentilis777@gmail.com
Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae) of the Crimea
(Ukraine). Krivosheyev R. E. – Twenty species of the Pselaphinae (Coleoptera, Staphylinidae) are
recorded for the first time from Crimean Peninsula. Bryaxis moczarskii (Winkler, 1911) is redescribed
and illustrated for the first time after its original description. Brachygluta guillemardi (Saulcy, 1876),
Bryaxis longulus (Kiesenwetter, 1849) and Claviger colchicus Motschulsky, 1837 are recorded for the
first time from Ukraine. In total, 34 species of ten genera and one unrecognizable nominal species are
recorded from Crimea.
Ke y wo r d s: Coleoptera, Staphylinidae, Pselaphinae, Crimea, Ukraine.
Îáçîð æóêîâ-îùóïíèêîâ (Coleoptera, Staphylinidae, Pselaphinae) Êðûìà (Óêðàèíà).
Êðèâîøååâ Ð. Å. – Âïåðâûå äëÿ Êðûìñêîãî ïîëóîñòðîâà óêàçàíû 20 âèäîâ îùóïíèêîâ.
Ïðèâåäåíî èëëþñòðèðîâàííîå ïåðåîïèñàíèå Bryaxis moczarskii (Winkler, 1911). Brachygluta guil-
lemardi (Saulcy, 1876), Bryaxis longulus (Kiesenwetter, 1849) è Claviger colchicus Motschulsky, 1837
îòìå÷åíû âïåðâûå â ôàóíå Óêðàèíû.  öåëîì èç Êðûìà ïðèâåäåíû 34 âèäà èç 10 ðîäîâ è îäèí
íå ïîäëåæàùèé èäåíòèôèêàöèè íîìèíàëüíûé âèä.
Êëþ÷åâûå ñ ëîâ à: Coleoptera, Staphylinidae, Pselaphinae, Êðûì, Óêðàèíà.
Introduction
The subfamily Pselaphinae included 70 species known hitherto from Ukraine and only 11 from Crimean
Peninsula. In Ukraine, the pselaphines were subject to special studies only in Lviv, Ternopil, Kyiv and
Transcarpathian Regions. All other territories were covered by a few publications representing mainly check-
lists or local records; there are no records of pselaphines at all from many regions of Ukraine. Many descrip-
tions and keys were incomplete, and most papers about Ukrainian pselaphines written a time ago (Nowicki,
1873, Lindemann, 1871; Weise, 1875; Jacobson, 1910; Roubal, 1930; Kryzhanovsky, 1965), except one
recent paper on transcarpathian beetles.
However, there are only 4 publications about Crimean pselaphines, including descriptions of Bryaxis
moczarskii Winkler, 1911, and with Euplectus frater Besuchet, 1964 and Tychus anatolicus Besuchet, 1964.
The other two papers are the brief reviews of the beetles from the south of Ukraine (with 3 species of pse-
laphines, Brachygluta furcata (Motschulsky, 1835), B. foveola (C. G. Thomson, 1859), B. helferi (Schmidt-
Goebel, 1836) recorded by Blinstein (1989) and the list of Pselaphidae from Russia and former USSR coun-
tries (Kurbatov, 2007), with additional 6 species known to Crimea. One nominal species, Batrisus tauricus
Motschulsky, 1851, is unrecognizable: its type material is missing and original description is incomplete and
uncertain. Thus, 11 species had been recorded by far from Crimean Peninsula before this study.
Crimean Peninsula represents diverse types of landscapes, from alkali soils and dry steppes of its north-
ern part to the mountain beech forests of Crimean Mountains and Mediterranean bush thickets. Such vari-
ety of biotopes suggests possible existence of quite diverse fauna, either common with Caucasian or
Mediterranean regions or endemic to Crimea.
Material and methods
Specimens were collected by sifting from decaying leaf litter of the beech and oak forest floor under
logs and near the large stones, killed in ethanol and then mounted. Dissections were made in water after
Vestnik zoologii, 47(4): 299—307, 2013
DOI 10.2478/vzoo-2013-0031
Unauthenticated
Download Date | 12/13/16 7:53 PM
maceration in KOH. Genitalia were placed in microvials with glycerin and pinned underneath the specimens.
Also a number of specimens deposited in the State Nature Museum of V. M. Karazin National University of
Kharkiv collection, collection of V. Lazorko at the Schmalhausen Institute of Zoology, or provided by
S. A. Kurbatov from Moscow State University, were examined. Pictures were taken with Leica M165C binoc-
ular microscope. The specimens are deposited in the collections of I. I. Schmalhausen Institute of Zoology,
Kyiv (SIZK), National Scientific Natural History Museum, Kyiv (NSNM), Museum of Nature of
V. M. Karazin National University of Kharkiv (MNKK), Donetsk National University (DNU), Zoological
Museum, University of Moscow (ZMUM) and Museum of Geneva.
Euplectus validus Besuchet, 1958
Besuchet, 1974: 318; Löbl, 2009: 6.
Ma t e r i a l. Big Canyon Reserve, under the maple bark, 6.05.1999, 1 ex. (Drogvalenko) (MNKK).
First record from the Crimea.
R ema r k s. E. validus can be easily differentiated from other species of the genus
Euplectus by having only two fossae on the elytra instead of four in other species.
D i s t r i b u t i o n i n Eu r o p e. Italy, Switzerland (Loebl, 2004).
Euplectus duponti Aubé, 1833
Besuchet, 1974: 319; Löbl, Besuchet, 2004: 284; Löbl, 2009: 4.
Ma t e r i a l. Yalta Highland Forest Reserve, near Yalta, under the bark of dead pine tree, 10.09.1999,
1 ex. (Drogvalenko) (MNKK). First record from Crimea.
D i s t r i b u t i o n i n Eu r o p e. West, South-West and Central Europe (Loebl,
2004).
Euplectus frater Besuchet, 1964
Besuchet, 1964: 415; 1974: 322; Löbl, Besuchet, 2004: 284; Löbl, 2009: 4.
Ma t e r i a l. The holotype was found in “Iaila-Dagh” (Winkler), and one of the paratypes was collect-
ed in “Baidar-Thor” (Knirsch) (Besuchet, 1974).
D i s t r i b u t i o n i n Eu r o p e. Ukraine, South and South-East Europe (Loebl,
2004).
Batrisodes venustus (Reichenbach, 1816)
Besuchet, 1974: 332; Löbl, Besuchet, 2004: 278; Löbl, 2009: 12.
Ma t e r i a l. Yalta Highland Forest Reserve, near the Uchan-Su waterfall, under the bark of pine tree
log, 16.05.2000, 2 ex. (Yunakov) (MNKK).
D i s t r i b u t i o n i n Eu r o p e. The whole Europe, except the South (Loebl,
2004). First record from Crimea.
Batrisodes hubenthali Reitter, 1913
Besuchet, 1974: 332; Löbl, Besuchet, 2004: 277; Löbl, 2009: 12.
Ma t e r i a l. South slope of Ai-Petri Yaila, vicinity of Uchan-Su waterfall, under pine tree bark,
17.05.2001, 2 ex. (Drogvalenko); Alushta District, vicinity of Izobilnoe Village, oak and beech forest, under
the oak bark, 9.05.2001, 1 ex. (Nadein) (MNKK). First record from Crimea.
D i s t r i b u t i o n i n Eu r o p e. Central, North Europe, some countries of Balkan
Peninsula, Italy, Ukraine (Loebl, 2004).
Batrisodes adnexus (Hampe, 1863)
Besuchet, 1974: 332; Löbl, Besuchet, 2004: 277; Löbl, 2009: 10.
Ma t e r i a l. Alushta District, vicinity of Izobilnoe Village, oak and beech forest, in the oak timber,
9.05.2001, 1 ex. (Nadein) (MNKK). First record from Crimea.
300 R. E. Krivosheyev
Unauthenticated
Download Date | 12/13/16 7:53 PM
Di s t r i b u t i o n i n Eu r o p e. The whole Europe except Scandinavia, Greece,
Spain, Great Britain and Baltic (Loebl, 2004).
Batrisodes oculatus (Aubé, 1833)
Besuchet, 1974: 333; Löbl, Besuchet, 2004: 277; Löbl, 2009: 12.
Ma t e r i a l. Vicinity of Sevastopol, under stone, with ants, 5.09.1920, 1 {, 1 } (Kizeritsky) (ZMUM).
First record from Crimea.
D i s t r i b u t i o n i n Eu r o p e. Mainly in the West of Europe, some of the Balcan
countries, and Finland, but present in Ukraine (Loebl, 2004).
Brachygluta retowskii (Simon, 1883)
Jakobson, 1910: 578; Besuchet, 1974: 353; Löbl, Besuchet, 2004: 298; Löbl, 2009: 14.
Ma t e r i a l. 10 km SE Eupatoria, Pribrezhnaya station, 4—6.05.1998, 1 ex. (Gorbunov) (MNKK);
Saki, on boggy places in the coast, 17.09.1954, 5 ex. (Bogachev) (MNKK).
D i s t r i b u t i o n i n Eu r o p e. Ukraine, South of Russia, Moldova, Romania,
Hungary, Slovakia and Austria (Loebl, 2004). Recorded from Crimea by Jakobson
(1910).
Brachygluta foveola (C. G. Thomson, 1859)
Blinstein, 1989: 61; Löbl, Besuchet, 2004: 297.
No material from Crimea was available in this study.
D i s t r i b u t i o n i n Eu r o p e. Ukraine, South of Russia, Great Britain, South of
Europe (Loebl, 2004).
Brachygluta xanthoptera Reichenbach, 1816
Besuchet, 1974: 354; Löbl, Besuchet, 2004: 299; Löbl, 2009: 14.
Ma t e r i a l. “Taur.”, 1 ex. (Hochhuth) (NSNM).
D i s t r i b u t i o n i n Eu r o p e. West, South and South-West of Europe, Ukraine
(Loebl, 2004). First record from Crimea.
Brachygluta helferi (Schmidt-Goebel, 1836)
Besuchet, 1974: 355; Blinstein, 1989: 59; Löbl, Besuchet, 2004: 298; Löbl, 2009: 12.
No material from Crimea was available in this study.
D i s t r i b u t i o n i n Eu r o p e. Almost whole Europe (Loebl, 2004).
Brachygluta guillemardi (Saulcy, 1876)
Besuchet, 1974: 354; Löbl, Besuchet, 2004: 297; Löbl, 2009: 12.
Ma t e r i a l. “Yalta 81”, 1 ex. (NSNM).
D i s t r i b u t i o n i n Eu r o p e. Slovenia, Croatia, Serbia, Bosnia, Italy, Spain,
France and Switzerland (Loebl, 2004). First record from Ukraine and Crimea.
Brachygluta klimschi Holdhaus, 1902
Besuchet, 1974: 355; Löbl, Besuchet, 2004: 298; Löbl, 2009: 14.
Ma t e r i a l. Simpheropol, Anatra, 28.02.2001, 69 ex. (Mosyakin) (SIZK).
301Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae)...
Unauthenticated
Download Date | 12/13/16 7:53 PM
Di s t r i b u t i o n i n Eu r o p e. Great Britain, Germany, Switzerland, Austria,
Slovenia, Italy (Loebl, 2004). First record from Crimea.
Brachygluta furcata (Motschulsky, 1835)
Blinstein, 1989: 61; Löbl, Besuchet, 2004: 297.
No material from Crimea was available in this study.
D i s t r i b u t i o n i n Eu r o p e. Ukraine, Italy, Greece, Albania, Macedonia,
South of Russia (Loebl, 2004).
Brachygluta simplicior Raffray, 1904
Besuchet, 1974: 356; Löbl, Besuchet, 2004: 299; Löbl, 2009: 14.
Ma t e r i a l. Simpheropol, Anatra, 28.02.2001, 1 ex. (Mosyakin) (SIZK).
D i s t r i b u t i o n i n Eu r o p e. South and West of Europe (Alonso-Zarazaga,
2004). First record from Crimea.
Brachygluta sinuata (Aubé, 1833)
Besuchet, 1974: 356; Löbl, Besuchet, 2004: 299; Löbl, 2009: 14.
Ma t e r i a l. “Simpheropol, Dzhamman”, 21.04.1951, 2 ex. (no collector) (MNKK).
D i s t r i b u t i o n i n Eu r o p e. Almost the whole Europe (Loebl, 2004). First
record from Crimea.
Brachygluta haematica (Reichenbach, 1816)
Löbl, Besuchet, 2004: 297; Löbl, 2009: 12.
Ma t e r i a l. Simpheropol Province, Tauric District, Dzhamman, 2 ex. (Kakhovsky) (MNKK).
D i s t r i b u t i o n i n Eu r o p e. Most part of Europe except Finland, Baltic and
Spain (Loebl, 2004). First record from Crimea.
Brachygluta nodosa (Motschulsky, 1835)
Löbl, Besuchet, 2004: 298.
Ma t e r i a l. Crimea, Bilohorsky District, near Topolevki Village, at a stream, in a rotten poplar,
31.07.1979, 1 ex. (Kurbatov, ZMUM).
D i s t r i b u t i o n i n Eu r o p e. Azerbaijan, Armenia, Georgia, South of Russia,
Ukraine (Crimea) (Loebl, 2004).
Reichenbachia juncorum Leach, 1817
Besuchet, 1974: 356; Löbl, Besuchet, 2004: 300; Löbl, 2009: 14.
Ma t e r i a l. “Taur.”, 1 ex. (Hochhuth) (NSNM).
D i s t r i b u t i o n i n Eu r o p e. North, Central and West (Loebl, 2004). First
record from Crimea.
Rybaxis longicornis (Leach, 1817)
Besuchet, 1974: 351; Löbl, Besuchet, 2004: 300; Löbl, 2009: 14.
Ma t e r i a l. “Taur.”, 2 ex. (Hochhuth) (NSNM).
D i s t r i b u t i o n i n Eu r o p e. Most of Europe except Norway and Baltic (Loebl,
2004). First record from Crimea.
302 R. E. Krivosheyev
Unauthenticated
Download Date | 12/13/16 7:53 PM
Bryaxis kruegeri (Machulka, 1932)
Besuchet, 1974: 342; Löbl, Besuchet, 2004: 306; Löbl, 2009: 18.
Ma t e r i a l. Nikitskaya Yaila, comb, in litter, 13.04.2001, 1 { (Yeremenko) (MNKK).
D i s t r i b u t i o n i n Eu r o p e. Italy, Switzerland, Austria, Croatia (Loebl, 2004).
First record from Crimea.
Bryaxis cateniger Krauss, 1899
Besuchet, 1974: 342; Löbl, Besuchet, 2004: 304; Löbl, 2009: 16.
Ma t e r i a l. Bilohorsky District, slope of mt. Karabi-Yaila, Su-At tract, beech forest, in a beech hole,
44°50'46'' N, 34°28'27'' E, 29.07.2012, 2 { (Krivosheyev) (SIZK); SW of Perevalnoe Village, Yegerlyk-Su,
Dolgorukovskaya Yaila, 18.06.2002, 2 {, 1 }; N-W Yalta, N slope of mt. Mogabi, h = 500 m, mixed for-
est, in litter, 23.05.2001, 1 { (Yunakov) (NMKK).
D i s t r i b u t i o n i n Eu r o p e. Italy, Austria, Slovenia (Alonso-Zarazaga, 2004).
First record from Crimea.
Bryaxis longulus (Kiesenwetter, 1849)
Besuchet, 1974: 342; Löbl, Besuchet, 2004: 307; Löbl, 2009: 18.
Ma t e r i a l. Zybino, Bilohorsky District, gardens, 9.05.1951, 2 { (Bogachev) (MNKK).
D i s t r i b u t i o n i n Eu r o p e. Italy, Austria, Slovenia, Croatia (Loebl, 2004).
First record from Ukraine and Crimea.
Bryaxis bulbifer (Reichenbach, 1816)
Besuchet, 1974: 348; Löbl, Besuchet, 2004: 304; Löbl, 2009: 16.
Ma t e r i a l. Yalta Highland Forest Nature Reserve, vicinity of Yalta, on lake bank, 8.05.1999, 8 {,
6 } (Yunakov) (MNKK); Simpheropol District, 2 km N of Privolnoe, at the foot of Mt. Demerdzhi, slopes
with beech and oak forest, in leaf litter near and under oak and beech logs along Kurliuk-Su River,
44°48'18'' N, 34°20'33'' E, 5—7.11.2012, 1 { (Krivosheyev) (SIZK).
D i s t r i b u t i o n i n Eu r o p e. All Europe, except Greece, Romania and Portugal
(Loebl, 2004). First record from Crimea.
Bryaxis moczarskii (Winkler, 1911) (fig. 1, 2)
Winkler, 1911: 7; Löbl, Besuchet, 2004: 307.
Ma t e r i a l. Simpheropol Region, 2 km N of Privolnoe, at the foot of Mt. Demerdzhi, slopes with
beech and oak forest, in leaf litter near and under oak and beech logs along Kurliuk-Su River, 44°48'18'' N,
34°20'33'' E, 5—7.11.2012, 5 {, 3 } (Krivosheyev) (SIZK); Alushta Region, mt. Demerdzhi slope, Khapkhal
Ravine, beech forest, under beeches near Ulu-Uzen’ River, along the forest trail, sifted from forest litter,
44°48.2' N 34°26.5' E, 2—3.05.2011, 3 {, 1 } (Krivosheyev) (SIZK); Burulcha tract, h = 978 m, beech for-
est, in litter, 23.06.2002, 1 {; Su-At River, h = 820 m, slope of the mountain, in litter, 23.06.2002, 1 {;
Chatyr-Dag, near the Bim-Bash-Koba cave, beech forest, in litter, 21.06.2002, 8 {, 6 } (Yunakov); Great
Canyon Reserve: beech forest, under the stone, 4.05.1999, 1 { (Drogvalenko); idem, under the stone in mice
holes, 6.05.1999, 2 {; idem, Tea House tract, beech forest, in litter, 22.05.2000, 4 {; Yalta Mountain-Forest
Reserve, near Yalta, on the lake bank, 8.05.1999, 2 } (Yunakov); Perevalnoe, near Yegerlyk-Su River, h =
600 m, beech forest, in litter, 21.06.2002, 1 { (Drogvalenko) (MNKK).
R ed e s c r i p t i o n. Male. Body light brown, unicolour, body length 2.2 mm, max-
imum width of elytron 0.8 mm (fig. 1, 1). Head 1.3 times as long as wide, almost
smooth and shining, slightly narrowed anteriorly. Frons with narrow, elongate excava-
tion between antennal tubercles; excavation almost reaching middle of head, bordered
with two lateral carinae. Median longitudinal occipital carina well defined. Eye well
visible dorsally, composed of 12 facets, vertexal foveae well defined, in line of eyes.
Maxillary palp long, segment I pedunculate, segment II very small, both non-granu-
lated by sight, segment III 0.3 mm, not very long, twice as short as head, densely
punctate and covered with short setae. Antenna short, slightly exceeding base of
303Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae)...
Unauthenticated
Download Date | 12/13/16 7:53 PM
elytron, scape 1.3 times as long as wide and 2 times as long as pedicel; pedicel almost
as long as wide, subspherical, antennomere III twice as long as wide, antennomere IV
1.5 times as long as wide, antennomeres V—VIII subequal in length, IX—X subspher-
ical, terminal segment 1.9 times as long as wide, acute at apex. Scape rounded rect-
angular, with well defined trapezium-like secretory nodule at inner side, ending with
small disk; pedicel oval, with small carina (fig. 1, 2). Pronotum gibbous, smooth,
glossy, covered with long, sparse setae, about as long as head and 1.25 times as wide
304 R. E. Krivosheyev
Fig. 1. Bryaxis moczarskii: 1 – habitus (dorsal view); 2 – the first three antennal segments; 3 – pronotum
and elytra, dorsal view.
Ðèñ. 1. Bryaxis moczarskii: 1 – îáùèé âèä ñâåðõó; 2 – òðè ïåðâûõ ñåãìåíòà àíòåíí; 3 – ïåðåäíåñïèí-
êà è íàäêðûëüÿ, ñâåðõó.
Fig. 2. Bryaxis moczarskii, aedeagus.
Ðèñ. 2. Bryaxis moczarskii, ýäåàãóñ.
as long (fig. 1, 3). Elytr on at suture
about 1.75 times as long as pronotum,
thinly punctated, also covered with long
setae (fig. 1, 3). Legs massive, all femo-
ra with noticeable tubercle close to
apex, tibia I with large dent in its last
third, tibia II simple, thin, tibia III with
shallow deepening in apical third, with
small dent on apex (fig. 1,1). Aedeagus
elongate, not very solid, thin. Inner
sack oval, middle-sized, vanes narrow,
1.25 times as long as inner sack, narrow
in their last ¼ and extended to triangu-
lar top, with several small and 3 large
setae per vane. Parameres simple, non-
ramose, rodlike, pointed to apex, slen-
der (fig. 2). Female with simple femora
and tibiae, elongate and simple scape
twice as long as wide, oval and simple
pedicel without carina.
D i s t r i b u t i o n i n E u r o p e.
Ukraine (Crimea) (Loebl, 2004).
Unauthenticated
Download Date | 12/13/16 7:53 PM
Tychus anatolicus Besuchet, 1964
Besuchet, 1964: 429; Löbl, Besuchet, 2004: 318
Ma t e r i a l. South Krimea, Angarsky pass, 15.08.1976, 1 {, 8 } (Nikitsky) (ZMUM); idem, 1 {
(Belov) (ZMUM); idem, 1 } (Nikitsky) (ZMUM).
D i s t r i b u t i o n i n Eu r op e. Ukraine, Romania, Greece (Loebl, 2004).
Tychus laminiger Besuchet, 1969
Besuchet, 1969: 409; Löbl, Besuchet, 2004: 319.
Ma t e r i a l. South Crimea, Nikita Botanical Garden, soil samples, 21.10.1968, 1 { (Arnoldi)
(ZMUM).
D i s t r i b u t i o n i n Eu r op e. Ukraine, Greece (Loebl, 2004).
Tychus niger (Paykull, 1800)
Jacobson, 1910: 583; Besuchet, 1974: 350; Löbl, Besuchet, 2004: 320; Löbl, 2009: 22.
Ma t e r i a l. Crimea, vicinity of Perevalnoe vil., 16.10.1988, 1 { (Nesterov, SIZK).
D i s t r i b u t i o n i n Eu r op e. Almost all Europe, except Spain, Portugal, Belarus
and Russia (Loebl, 2004).
Tychus tauricus Motschulsky, 1851
Motschulsky, 1851: 488; Besuchet, Sabella 1999: 243; Löbl, Besuchet, 2004: 281 (Tychus).
Ma t e r i a l. Absent in Motschulsky collection in the Zoological Museum, University of Moscow.
R ema r k s. Identity of the species is uncertain: original description is poor.
According to Besuchet, Sabella (1999), it was often considered a synonym of T.
dichrous Schmidt-Goebel, 1836, which is considered a synonym of T. niger. Formal
synonymization possibly needs designation of the neotype for Tychus tauricus
Motschulsky, 1851.
D i s t r i b u t i o n i n Eu r op e. Ukraine (Loebl, 2004).
Pselaphus heisei (Herbst, 1792)
Besuchet, 1974: 358; Löbl, Besuchet, 2004: 327; Löbl, 2009: 24.
Ma t e r i a l. Crimea, near Perevalnoe Village, 15.10.1981, 1 ex. (Nesterov) (SIZK); Simpheropol,
Salgirka, park, 5.04.1957, 1 ex. (Bogachev) (SIZK).
D i s t r i b u t i o n i n Eu r op e. Almost all Europe, except Italy, Portugal and
Greece (Loebl, 2004). First record from Crimea.
Ctenistes palpalis (Reichenbach, 1816)
Besuchet, 1974: 360; Löbl, Besuchet, 2004: 322; Löbl, 2009: 24.
Ma t e r i a l. Simpheropol District, Dubki, 24.09.1909, 1 ex. (Kakhovsky) (MNKK); “Taur.”, 7 ex.
(Hochhuth) (NSNM); Simpheropol, dry limestone slopes, 26.10.1956, 1 ex. (Bogachev) (SIZK).
D i s t r i b u t i o n i n Eu r op e. All Europe, except North, Great Britain and
Poland (Loebl, 2004). First record from Crimea.
Claviger colchicus Motschulsky, 1837 (fig. 3)
Löbl, Besuchet, 2004: 281.
Ma t e r i a l. Chatyr-Dag, S slope, h = 1200, under a stone, 14.05.2006, 3 ex. (V. Martynov) (DNU);
Karabi-Yaila, under a stone, 8.05.2006, 1 ex. (Gorbunov) (MNKK).
D i s t r i b u t i o n i n Eu r op e. Azerbaijan, Armenia, Georgia, South of Russia
(Loebl, 2004). First record from Crimea and Ukraine.
305Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae)...
Unauthenticated
Download Date | 12/13/16 7:53 PM
Species incertae sedis
Batrisus tauricus Motschulsky, 1851
Motschulsky, 1851: 488; Löbl, Besuchet, 2004: 281.
Ma t e r i a l. Type specimens are missing; the original description is incomplete, and this species is
unrecognizable.
D i s t r i b u t i o n i n Eu r op e. Ukraine (Crimea) (Motschulsky, 1851).
Conclusion
Thirty-four species of ten pselaphine genera known to occur in Crimean Peninsula
are listed, including data on distribution and mention in the most important taxonomic
works. Three of them, Brachygluta guillemardi (Saulcy, 1876), Bryaxis longulus
(Kiesenwetter, 1849) and Claviger colchicus Motschulsky, 1837, are recorded for the first
time from Ukraine, and 21 species from Crimea. Two species (Bryaxis moczarskii
(Winkler, 1911) and Claviger colchicus Motschulsky, 1837) are illustrated for the first time.
The author thanks A. A. Drogvalenko (Kharkiv) and V. V. Martynov (Donetsk) for the possibility to
work at the collection of the State Museum of Nature of V. M. Karazin National University and Donetsk
National University, S. A. Kurbatov (Moscow) and S. A. Mosyakin (Simpheropol) for providing material
from Crimea. V. A. Korneyev (Kyiv) assisted me with preparation of illustrations, kindly read an early ver-
sion of this manuscript and gave valuable comments on the description and corrections of its English text. I
appreciate the work of S. A. Kurbatov and R. Bekchiev (Sophia), who kindly reviewed this paper and gave
numerous recommendations how to improve it.
References
Besuchet C. Psélaphides paléarctiques. Espèces nouvelles et notes synonimiques. II. Coleoptera // Revue
Suisse de Zoologie. – 1964. – 71. – P. 411—443.
Besuchet C. Psélaphides paléarctiques. Espèces nouvelles et notes synonimiques. III. Coleoptera // Revue
Suisse de Zoologie. – 1969. – 76. – P. 397—420.
306 R. E. Krivosheyev
Fig. 3. Claviger colchicus:1, 2 – habitus (dorsal view).
Ðèñ. 3. Claviger colchicus:1, 2 – îáùèé âèä ñâåðõó.
Unauthenticated
Download Date | 12/13/16 7:53 PM
Besuchet C. Pselaphidae // Die Käfer Mitteleuropas. Band 2. Staphylinidae II (Hypocyptinae and
Aleocharinae). Pselaphidae / Eds H. Freude, K. W. Harde, G. A. Löhse. – Krefeld : Goecke & Evers,
1974. – S. 305—362.
Besuchet C., Sabella G. Nuove specie di Tychus della regione paleartica (Coleoptera, Staphylinidae,
Pselaphinae) // Mitteilungen der Schweizerischen entomologischen Gesellschaft. – 1999. –72. –
P. 235—258.
Blinstein S. Ya. On the coleopterans of the South of Ukraine // Ecology and taxonomy of insects of
Ukraine. – Kiev : Vyshcha shkola, 1989. – P. 59—64. – Russian : Áëèíøòåéí Ñ. ß. Ê èçó÷åíèþ
æåñòêîêðûëûõ þãà Óêðàèíû.
Jacobson G. G. 12. Family Clavigeridae (Pselaphidae) // Beetles of Russia and West Europe. – S-Petersburg :
F. Devrien, 1910. – Vol. 8 – P. 569—588 + Tab. 62—68, 70. – Russian : ßêîáñîí Ã. Ã. 12. Ñåìåéñòâî
Clavigeridae (Pselaphidae). Îùóïíèêè.
Kryzhanovsky O. L. 17. Fam. Pselaphidae – short-winged mold beetles // Keys to insects of of European
Territory of USSR. Coleoptera and Strepsiptera. – Moscow ; Leningrad : Nauka, 1965. – Vol. 2. –
P. 156—158. – Russian : Êðûæàíîâñêèé Î. Ë. 17. Ñåì. Pselaphidae – îùóïíèêè.
Kurbatov S. A. List of the beetles (Pselaphidae) of Russia and countries of the former USSR. – http: //
www.zin.ru/Animalia/Coleoptera/rus/incops1.htm. – 2007. – Accessed on January 14, 2013. –
Russian : Êóðáàòîâ Ñ. À. Ñïèñîê îùóïíèêîâ (Pselaphidae) Ðîññèè è ñòðàí áûâøåãî ÑÑÑÐ.
Lindemann K. A review of geographic distribution of beetles in Russian Empire. Part 1. Introduction, Fore
word. Northern, Moscowian and Turanian Provinces // Proceedings of Russian Entomological
Society. – 1871. – Vol. 6. – P. 41—366. – Russian : Ëèíäåìàí Ê. Îáçîð ãåîãðàôè÷åñêîãî ðàñïðî-
ñòðàíåíèÿ æóêîâ â Ðîññèéñêîé Èìïåðèè. ×àñòü 1. Ââåäåíèå, ïðåäèñëîâèå. Ñåâåðíàÿ,
Ìîñêîâñêàÿ è Òóðàíñêàÿ ïðîâèíöèè.
Löbl I. Coleoptera Staphylinidae (Dasycerinae, Pselaphinae) // Folia Heyrovskyana, series B. Icones
Insectorum Europae Centralis. – Zlín : Kabourek Publishing, 2009. – 28 p.
Löbl I., Besuchet C. Pselaphinae // Catalogue of Palearctic Coleoptera.– Stenstrup : Apollo Books, 2004. –
Vol. 2. – P. 272—329.
Loebl I. Pselaphinae // Coleoptera, Beetles. – Fauna Europaea, version 1.1 / Ed. M. A.Alonso-Zarazaga.–
2004. – Acessed on 1.12.2012 . – http: // www.faunaeur.org.
Motschulsky V. Enumération des nouvelles espèces de coléoptères de son dernier voyage // Bulletin de la
Société Impériale des Naturalistes du Moscou. – 1851. – 24, N 4. – P. 479—511.
Nowicki M. Pselaphidae. Clavigeridae // Beiträge zur Insektenfauna Galiziens. – Krakau : Jagellonische
Universitäts-Buchdrucketei, 1873.– S. 20.
Reitter E. Fauna Germanica // Die Käfer des Deutschen Reiches. – Stuttgart : K. G. Lutz Verl, 1909. –
Bd. 2. – 392 S.
Roubal J. Katalog Coleopter (brouků) Slovenska a Podkarpatska Rus. T. 1. – Praha : Nakladatelstvi Orbis,
1930. – 527 s.
Weise J. Coleoptrologische Ergebnisse einer Bereisung der Czernahora // Verh. Naturforsch. Vereines in
Brünn. – 1875. – 14. – S. 85—114.
Received 18 January 2013
Accepted 20 May 2013
307Review of the Short-Winged Mold Beetles (Coleoptera, Staphylinidae, Pselaphinae)...
Unauthenticated
Download Date | 12/13/16 7:53 PM
|