A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine
A new fossil species Perca neopleistocenica Kovalchuk, sp. n. from Southern Ukraine is described. It differs from other species of the genus by width and configuration of praeoperculum, and also by quantity of cogs on the posterior edge of os praeoperculare.
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Інститут зоології ім. І.І. Шмальгаузена НАН України
2013
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| Цитувати: | A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine / О.М. Kovalchuk // Вестник зоологии. — 2013. — Т. 47, № 4. — С. 377—380. — Бібліогр.: 19 назв. — англ. |
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Digital Library of Periodicals of National Academy of Sciences of Ukraine| id |
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irk-123456789-1098012016-12-15T03:02:49Z A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine Kovalchuk, О.М. Краткие сообщения A new fossil species Perca neopleistocenica Kovalchuk, sp. n. from Southern Ukraine is described. It differs from other species of the genus by width and configuration of praeoperculum, and also by quantity of cogs on the posterior edge of os praeoperculare. Описан новый ископаемый вид Perca neopleistocenica Kovalchuk, sp. n. из южной Украины. Он отличается от других видов рода шириной и конфигурацией предкрышки, а также количеством зубцов на заднем крае os praeoperculare. 2013 Article A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine / О.М. Kovalchuk // Вестник зоологии. — 2013. — Т. 47, № 4. — С. 377—380. — Бібліогр.: 19 назв. — англ. 0084-5604 DOI 10.2478/vzoo-2013-0041 http://dspace.nbuv.gov.ua/handle/123456789/109801 567(477.64) en Вестник зоологии Інститут зоології ім. І.І. Шмальгаузена НАН України |
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Краткие сообщения Краткие сообщения |
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Краткие сообщения Краткие сообщения Kovalchuk, О.М. A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine Вестник зоологии |
| description |
A new fossil species Perca neopleistocenica Kovalchuk, sp. n. from Southern Ukraine is described. It differs from other species of the genus by width and configuration of praeoperculum, and also by quantity of cogs on the posterior edge of os praeoperculare. |
| format |
Article |
| author |
Kovalchuk, О.М. |
| author_facet |
Kovalchuk, О.М. |
| author_sort |
Kovalchuk, О.М. |
| title |
A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine |
| title_short |
A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine |
| title_full |
A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine |
| title_fullStr |
A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine |
| title_full_unstemmed |
A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine |
| title_sort |
new fossil perca fish (teleostei, percidae) from the neopleistocenian sediments of southern ukraine |
| publisher |
Інститут зоології ім. І.І. Шмальгаузена НАН України |
| publishDate |
2013 |
| topic_facet |
Краткие сообщения |
| url |
http://dspace.nbuv.gov.ua/handle/123456789/109801 |
| citation_txt |
A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine / О.М. Kovalchuk // Вестник зоологии. — 2013. — Т. 47, № 4. — С. 377—380. — Бібліогр.: 19 назв. — англ. |
| series |
Вестник зоологии |
| work_keys_str_mv |
AT kovalchukom anewfossilpercafishteleosteipercidaefromtheneopleistoceniansedimentsofsouthernukraine AT kovalchukom newfossilpercafishteleosteipercidaefromtheneopleistoceniansedimentsofsouthernukraine |
| first_indexed |
2025-07-07T23:40:38Z |
| last_indexed |
2025-07-07T23:40:38Z |
| _version_ |
1837033470797283328 |
| fulltext |
UDC 567(477.64)
A NEW FOSSIL PERCA FISH (TELEOSTEI, PERCIDAE)
FROM THE NEOPLEISTOCENIAN SEDIMENTS
OF SOUTHERN UKRAINE
Î. Ì. Kovalchuk
National Museum of Natural History NAS of Ukraine,
vul. B. Khmelnitskogo, 15, Kyiv, 01601 Ukraine
E-mail: Biologiest@ukr.net
A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian Sediments of Southern Ukraine.
Kovalchuk Î. Ì. – A new fossil species Perca neopleistocenica Kovalchuk, sp. n. from Southern
Ukraine is described. It differs from other species of the genus by width and configuration of praeop-
erculum, and also by quantity of cogs on the posterior edge of os praeoperculare.
Ke y wo r d s: Perca, new species, os praeoperculare, alluvium, beginning of the Middle Neopleisto -
cene, Zaporozhye region, Ukraine.
Íîâûé èñêîïàåìûé âèä ðîäà Perca (Teleostei, Percidae) èç íåîïëåéñòîöåíîâûõ îòëîæåíèé íà þãå
Óêðàèíû. Êîâàëü÷óê À. Í. – Îïèñàí íîâûé èñêîïàåìûé âèä Perca neopleistocenica Kovalchuk,
sp. n. èç þæíîé Óêðàèíû. Îí îòëè÷àåòñÿ îò äðóãèõ âèäîâ ðîäà øèðèíîé è êîíôèãóðàöèåé ïðåä-
êðûøêè, à òàêæå êîëè÷åñòâîì çóáöîâ íà çàäíåì êðàå os praeoperculare.
Êëþ÷åâûå ñ ëîâ à: Perca, íîâûé âèä, os praeoperculare, àëëþâèé, íà÷àëî ñðåäíåãî íåîïëåé-
ñòîöåíà, Çàïîðîæñêàÿ îáëàñòü, Óêðàèíà.
Introduction
New locality of the fossil fauna Lysa Gora was discovered in 2009 during the field paleozoological
investigations of expedition of the National Museum of the Natural History, National Academy of Sciences
of Ukraine. It is situated near Vasilyevka, Zaporozhye region (Southern Ukraine). This locality is a natural
outcrop (to 20 m high) of Miocenian and Pleistocenian sediments on the left bank of the Dnieper River.
Two horizons of sands and gravelites with numerous fossil remains of fishes and mammals were selected in
deposits of clays, limestones, loesses and fossil soils. Lysa Gora—1 is a former Neopleistocenian river avan-
delta.
Remains of perch from these sediments were found to be different from all other species of the genus
in a number of characters and described here as a new species. Determination of the fossil remains was real-
ized by comparison with collection of the recent and fossil material, and also with use of corresponding zoo-
logical (Shcherbukha, 1981) and paleontological works (Radu, 2005). Description of the new taxon was car-
ried out on the scheme, proposed by I. A. Korobkov for the fossils (Korobkov, 1971), on the basis of the
special work of J. Lepiksaar (1994) and principles of acting International Codex of Zoological Nomenclature
(International Code..., 2003).
Perca neopleistocenica Kovalchuk, sp. n.
Mat e r i a l. The holotype (fig. 1) is a near-complete left praeoperculum. It is deposited in the
Department of Vertebrate Paleozoology and Paleontological Museum, National Museum of the Natural
History (NMNH), National Academy of Sciences of Ukraine (Kyiv).
Referred material. One praeoperculum certainly belong to the new taxon. The attribution of a few
postcranial bones (vertebrae etc.) is less certain.
L o c a l i t y. Lysa Gora—1 (47°28.373“ N, 35°16.114“ E), left bank of the Dnieper
River, near Vasilyevka, Zaporozhye region, Southern Ukraine.
Geo l o g i c a l a g e . Beginning of the Middle Neopleistocene (Likhvin age,
Holstein, MQR 3, OIS 11).
Vestnik zoologii, 47(4): 377—380, 2013
DOI 10.2478/vzoo-2013-0041
Unauthenticated
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Dia gno s i s. New fossil Perca species is characterized by broadest praeoperculum
with smaller, frequent massive rounded cogs.
De s c r i p t i o n a nd c ompa r i s on. The maximum preserved praeopercular
length is about 1.1 cm (total length of the complete bone is near 4 cm). Considering a
directly proportional dependence between viscerocranial bones and total fish body
length (Lebedev, 1960), and also on the basis of comparison with praeoperculum of
recent Perca fluviatilis Linnaeus, 1758 it is safe to say, that total body length of the new
species is near 20 cm.
A surface of bone is raised, slightly bulging. Angulus superior is destroyed, angu-
lus anterior is partially broken. Pori canales lineae lateralis are well-preserved. Medial
part of the os praeoperculare is sharply defined, with developed relief. Praeopercular
limbus posterior is jagged, similar to that in all species of Perca Linnaeus, 1758.
Cogs of the os praeoperculare should be uniform in size, whereas Gymnocephalus
Bloch, 1793, the nearest to Perca genus of the family Percidae Cuvier, 1816 based on
mitochondrial DNA sequence (Song et al., 1998; Mamilov et al., 2003; Sloss et al.,
2004), has three long large cogs at the point of praeopercular angle. Contrary to the
praeoperculum of Perca fluviatilis Linnaeus, which has flat saw-like infrequent cogs,
praeopercular cogs of the new species are more frequent (cogs quantity proportion
between P. fluviatilis and P. neopleistocenica is 1 : 2), massive, adjoined face to face and
rounded in the cross section.
Judging from the preserved lamina praeopercularis, the angle among two ends of
bone is 120° (praeopercular angle of P. fluviatilis is 90°). Plate of the praeoperculum is
broader, its maximum width is 0.64 ñm.
E t ymo l o g y. The specific name is a Latin derivation and reflects the age of cor-
responding alluvial horizon.
Attendant material. Besides the perch remains, pharyngeal teeth of cyprinid fishes
(Cyprinus cf. carpio, Cyprinus sp., Carassius carassius, Carassius sp., Tinca sp., Rutilus
cf. frisii, Rutilus sp., Scardinius erythrophthalmus, Scardinius sp., Leuciscus sp.,
Chondrostoma cf. nasus, Chondrostoma sp.), lateral ethmoid of Cobitis sp., teeth and
bones of Silurus sp., Esox lucius, Esox sp., also some bone fragments of large and small
mammals were found in the same alluvial layer of Lysa Gora—1.
D i s t r i b u t i o n. ? Middle Neopleistocene of Southern Ukraine.
378 Î. Ì. Kovalchuk
Fig. 1. Perca neopleistocenica sp. n., left praeoperculum: 1 – anterior view; 2 – posterior view.
Ðèñ. 1. Perca neopleistocenica sp. n., ëåâûé praeoperculum: 1 – âèä cïåðåäè, 2 – âèä ñçaäè.
Unauthenticated
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Rema r k s . Perca neopleistocenica is described on the basis of fossil praeoperculum
collected from the Neopleistocenian locality Lysa Gora—1. The age of fossil material is
estimated to be 1,8 Ma. On the territory of Ukraine, fossil remains of European perch
are known from the Middle-Pliocenian deposits near Kamenskoye, Zaporozhye region
(Tarashchuk, 1962), from the Late Pliocene of Kairy, Kherson region and Nogaisk,
Zaporozhye region (Tarashchuk, 1962). Besides numerous bones of P. fluviatilis were
found in the Middle-Pleistocenian and Early-Holocenian deposits near Kaniv and Kyiv
(Shpet, 1949), on the territory of Palaeolithic site on the right bank of the Desna River
near Novgorod-Siverskiy (Nikolsky, 1952), in materials of the mound burier of
Õ—ÕI ñent. Shestovitsy (Chernigov region) and Slavic settlement of VIII—XIII cent. on
the bank of Vorskla River, Poltava region (Lebedev, 1960).
The recent percids are represented by a wide-variety of morphological forms. They
are the subject of numerous phylogenetic studies (Svetovidov, Dorofeeva, 1963;
Collette, 1963; Collette, Bănărescu, 1977; Song et al., 1998; Mamilov et al., 2003).
Twelve species of 5 genera (Stizostedion, Perca, Gymnocephalus, Zingel, Percarina) of
the family Percidae are presented in the modern Ukrainian ichthyofauna (Movchan,
2008—2009). More than 190 percid species are in the world fauna (Sloss et al., 2004),
which allow to assume the evolutional prosperity of this group of freshawater fishes.
Conclusions
Finding a new fossil Perca in Neopleistocenian deposits in Zaporozhye region adds
to our knowledge on the history of Ukrainian freshwater perciform fish. Fossil and
recent forms of the genus Perca are not fully studied till present (Shcherbukha, 1992).
The range of its affinity depends on their geological age, origin and environmental con-
ditions (Shcherbukha, 1993; Yakovlev, 1964).
Thus, fossil fishes appear to be useful subjects for discussing European palaeobio-
geography and freshwater environmental continuity. This encourages further develop-
ment of our ability to distinguish fossil fish species on the basis of disarticulated and
fragmentary osteological material, which is the case for most Ukrainian continental
material.
We extend our gratitude to Dr. E. Šwidnicka for access to the palaeontological collection in the
Department of Paleozoology (Zoological Institute, Wrocław University, Poland). We also thank
Prof. L. I. Rekovets for a giving possibility to take part in the field investigations on Lysa Gora locality
(2009—2011) and for his constant support. The drawings were prepared by M. V. Sinitsa (NMNH, Kyiv).
The manuscript was extensively reviewed by Dr. L. I. Klochko (Makarenko Sumy State Pedagogical
University, Sumy, Ukraine) whose kindness is greatly acknowledged.
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379A New Fossil Perca Fish (Teleostei, Percidae) from the Neopleistocenian...
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íîâîäíûõ ðûá.
Received 19 November 2012
Accepted 20 May 2013
380 Î. Ì. Kovalchuk
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