New nomenclatural combinations in Blitum L. and Kali Mill. (Chenopodiaceae): taxa occurring in Eastern Europe
П’ять нових номенклатурних комбінацій (чотири на рівні секції й одна для гібрида) запропоновані в роді Blitum L. та одна нова комбінація на рівні підвиду — в роді Kali Mill. (Chenopodiaceae). Стисло обґрунтована необхідність виділення цих родів зі складу Chenopodium L. і Salsola L. відповідно....
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irk-123456789-1746772021-01-28T01:26:50Z New nomenclatural combinations in Blitum L. and Kali Mill. (Chenopodiaceae): taxa occurring in Eastern Europe Mosyakin, S.L. Судинні рослини: систематика, географія, флора П’ять нових номенклатурних комбінацій (чотири на рівні секції й одна для гібрида) запропоновані в роді Blitum L. та одна нова комбінація на рівні підвиду — в роді Kali Mill. (Chenopodiaceae). Стисло обґрунтована необхідність виділення цих родів зі складу Chenopodium L. і Salsola L. відповідно. Пять новых номенклатурных комбинаций (четыре на уровне секции и одна для гибрида) предложены в роде Blitum L. и одна новая комбинация на подвидовом уровне — в роде Kali Mill. (Chenopodiaceae). Кратко обоснована необходимость выделения этих родов из состава, соответственно, Chenopodium L. и Salsola L. 2012 Article New nomenclatural combinations in Blitum L. and Kali Mill. (Chenopodiaceae): taxa occurring in Eastern Europe / S.L. Mosyakin // Український ботанічний журнал. — 2012. — Т. 69, № 3. — C. 393-396. — Бібліогр.: 19 назв. — англ. 0372-4123 http://dspace.nbuv.gov.ua/handle/123456789/174677 en Український ботанічний журнал Інститут ботаніки ім. М.Г. Холодного НАН України |
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Судинні рослини: систематика, географія, флора Судинні рослини: систематика, географія, флора |
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Судинні рослини: систематика, географія, флора Судинні рослини: систематика, географія, флора Mosyakin, S.L. New nomenclatural combinations in Blitum L. and Kali Mill. (Chenopodiaceae): taxa occurring in Eastern Europe Український ботанічний журнал |
| description |
П’ять нових номенклатурних комбінацій (чотири на рівні секції й одна для гібрида) запропоновані в роді Blitum L. та одна нова комбінація на рівні підвиду — в роді Kali Mill. (Chenopodiaceae). Стисло обґрунтована необхідність виділення цих родів зі складу Chenopodium L. і Salsola L. відповідно. |
| format |
Article |
| author |
Mosyakin, S.L. |
| author_facet |
Mosyakin, S.L. |
| author_sort |
Mosyakin, S.L. |
| title |
New nomenclatural combinations in Blitum L. and Kali Mill. (Chenopodiaceae): taxa occurring in Eastern Europe |
| title_short |
New nomenclatural combinations in Blitum L. and Kali Mill. (Chenopodiaceae): taxa occurring in Eastern Europe |
| title_full |
New nomenclatural combinations in Blitum L. and Kali Mill. (Chenopodiaceae): taxa occurring in Eastern Europe |
| title_fullStr |
New nomenclatural combinations in Blitum L. and Kali Mill. (Chenopodiaceae): taxa occurring in Eastern Europe |
| title_full_unstemmed |
New nomenclatural combinations in Blitum L. and Kali Mill. (Chenopodiaceae): taxa occurring in Eastern Europe |
| title_sort |
new nomenclatural combinations in blitum l. and kali mill. (chenopodiaceae): taxa occurring in eastern europe |
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Інститут ботаніки ім. М.Г. Холодного НАН України |
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2012 |
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Судинні рослини: систематика, географія, флора |
| url |
http://dspace.nbuv.gov.ua/handle/123456789/174677 |
| citation_txt |
New nomenclatural combinations in Blitum L. and Kali Mill. (Chenopodiaceae): taxa occurring in Eastern Europe / S.L. Mosyakin // Український ботанічний журнал. — 2012. — Т. 69, № 3. — C. 393-396. — Бібліогр.: 19 назв. — англ. |
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Український ботанічний журнал |
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2025-07-15T11:43:32Z |
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2025-07-15T11:43:32Z |
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1837713129796009984 |
| fulltext |
393ISSN 0372-4123. Укр. ботан. журн., 2012, т. 69, № 3
S.L. MOSYAKIN
M.G. Kholodny Institute of Botany, National Academy
of Sciences of Ukraine
2 Tereshchenkivska Street, Kyiv, 01601 Ukraine
inst@botany.kiev.ua
NEW NOMENCLATURAL COMBINATIONS
IN BLITUM L. AND KALI MILL.
(CHENOPODIACEAE): TAXA OCCURRING
IN EASTERN EUROPE
К e y wo r d s: new combinations, Blitum, Kali, Chenopodium, Salsola
© S.L. MOSYAKIN, 2012
Abstract. Five new combinations (four at the section level and one for
a hybrid) are validated in the genus Blitum L. and one new combination
at the subspecies level is made in the genus Kali Mill. (Chenopodiaceae).
Segregation of these genera from Chenopodium L. and Salsola L., re-
spectively, is briefly discussed.
Introduction
During the preparation of treatments of Chenopodium L. and related genera of Che no po-
dia ceae for the new checklist of the vascular flora of Eastern Europe being compiled at
the V.L. Ko ma rov Botanical Institute of the Russian Academy of Sciences, it has beco-
me evident that several new nomenclatural combinations are needed in these groups.
The new combinations validated below are partly based on our previous taxo-
nomic research (Mosyakin, 1993, Mosyakin, Clemants, 1997; Мосякін, 2002, 2003,
etc.), as well as on new results of molecular phylogenetic studies of Chenopodiaceae
(Akhani et al., 2007; Kadereit et al., 2003, 2005, 2010, etc.).
Discussion
Results of recent molecular studies confirmed the deep phylogenetic split between at
least three branches containing taxa previously traditionally placed in Chenopodium
sensu lato (Kadereit et al., 2003, 2005, 2010), roughly corresponding to commonly
recognized subgenera of Chenopodium (subgen. Ambrosia A.J. Scott, subgen. Blitum
(L.) I. Hiitonen, and subgen. Chenopodium), with addition of taxa from other tradi-
tionally recognized genera. In particular, these results confirmed the taxonomic and
nomenclatural decisions of S.L. Mosyakin and S.E. Clemants (2002, 2008), who
transferred to the expanded genus Dysphania R. Br. glandular-pubescent taxa previ-
ously placed in Chenopodium subgen. Ambrosia. The new circumscription of Dysphania
was used in treatments prepared for the Flora of North America north of Mexico
(Clemants, Mosyakin, 2003) and the Flora of China (Zhu Gelin et al., 2003), and now
is gaining wider recognition in other regional floras of the world and taxonomic and
floristic databases (Shepherd, Wilson, 2008, 2009; Verloove, Lambinon, 2006, etc.).
It seems now that the Linnaean genus Blitum L. has to be resurrected from its tem-
porary taxonomic oblivion. Consequently, new combinations will be needed for taxa
394 ISSN 0372-4123. Ukr. Botan. Journ., 2012, vol. 69, № 3
currently treated in Chenopodium, Monolepis Schrad., and probably some other groups,
for which no valid names are currently available in Blitum. The clade containing Blitum
also contains Spinacia L., which may cause serious nomenclatural problems if these two
simultaneously validated genera are considered congeneric. In my opinion, nomenclat-
ural conservation of Blitum against Spinacia would be desirable in the future, which
would minimize possible nomenclatural changes. Earlier we validated or re-circum-
scribed some infragenetic taxa within Chenopodium subgen. Blitum (Mosyakin, Clemants,
1997). Now these taxa have to be transferred to the genus Blitum.
In my treatments of Salsola L. sensu lato for the Flora of North America north of
Mexico (Mosyakin, 1996, 2003) I agreed with some other authors in recognizing that
Salsola, as traditionally circumscribed, is a non-monophyletic aggregate rather than a
natural genus. Recent molecular phylogenetic research (Kadereit et al., 2003; and espe-
cially Akhani et al., 2007) greatly contributed to a better understanding of relationships
between various representatives of Salsoloideae and Salsoleae. In particular, the genus
Kali Mill. was recognized to house taxa related to Kali soda Moench (= Salsola kali L.),
and several new combinations have been proposed (Akhani et al., 2007). However, one
infraspecific taxon occurring in Eastern Europe has not been transferred to the genus
Kali. The relevant nomenclatural transfer in Kali is also validated below.
Validation of new combinations
Here I validate new combinations only for taxa occurring in Eastern Europe, for their
subsequent inclusion in the mentioned annotated checklist of East European vascular
plants (in preparation). Other new names and combinations will be published later,
after further studies and proper discussion with colleagues.
Genus Blitum L.
Blitum L. sect. Blitum subsect. Foliosa (Kowal ex Mosyakin & Clemants) Mo sya-
kin, comb. nov.
Basionym: Chenopodium L. [subgen. Blitum (L.) I. Hiitonen Blitum (L.) Hook. f. in Benth. &
Hook. f.] subsect. Foliosa Kowal ex Mosyakin & Clemants, 1996, Novon 6(4): 398. — Typus:
Chenopodium foliosum Asch. (= Blitum virgatum L.).
— Chenopodium sect. Eublitum (Moq.) Aellen subsect. Foliosa Kowal, 1953, Monogr. Bot.
(War szawa) 1: 113, nom. inval. (descr. polon.).
This subsection includes taxa of the Blitum foliosum aggr., while B. capitatum L. (= Chenopodium
capitatum (L.) Ambrosi) should be placed in Blitum L. sect. Blitum subsect. Blitum (Chenopodium L.
subgen. Blitum (L.) I. Hiitonen sect. Blitum (L.) Hook. f. in Benth. & Hook. f. subsect. Capitata Kowal
ex Mosyakin & Clemants, 1996, Novon 6(4): 399. — Chenopodium sect. Eublitum (Moq.) Aellen sub-
sect. Capitata Kowal, 1953, Monogr. Bot. (Warszawa) 1: 113, nom. inval., descr. polon.).
Blitum L. sect. Degenia (Aellen) Mosyakin, comb. nov.
Basionym: Chenopodium sect. Degenia Aellen, 1926, Magyar Bot. Lapok, 25: 56. — Typus:
Chenopodium macrospermum Hook. f.
In the Eastern European flora this section is represented by B. chenopodioides L. (= Chenopodium
chenopodioides (L.) Aellen, C. botryodes Sm., C. crassifolium Hornem.).
Blitum L. sect. Pseudoblitum (Hook. f.) Mosyakin, comb. nov.
Basionym: Chenopodium L. sect. Pseudoblitum Hook. f. 1880, in Benth. et Hook. f., Gen. Pl. 3:
52. — Typus: Chenopodium rubrum L. (= Blitum rubrum (L.) Rchb.).
395ISSN 0372-4123. Укр. ботан. журн., 2012, т. 69, № 3
This section in Eastern Europe is represented by B. rubrum (L.) Rchb. (= Chenopodium rubrum L.).
Blitum L. sect. Glauca (Standl.) Mosyakin, comb. nov.
Basionym: Chenopodium L. [unranked] Glauca Standl. 1916, North Amer. Fl. 21: 29. — Typus:
Chenopodium glaucum L. (Blitum glaucum (L.) W.D.J. Koch).
— Chenopodium L. sect. Glauca Ignatov, 1988, Сосуд. раст. сов. Дальн. Вост. 3: 22.
This section includes B. glaucum (L.) W.D.J. Koch (= Chenopodium glaucum L.) that occurs in
Eastern Europe and has almost worldwide distribution, and some related taxa, usually treated as spe-
cies or infraspecific entities.
Blitum × schulzeanum (Murr) Mosyakin, comb. nov.
Basionym: Chenopodium × schulzeanum Murr, 1906, Allg. Bot. Zeitschr. 12: 110.
Blitum × schulzeanum is a hybrid between B. glaucum and B. rubrum; it is sporadically registered
in places of co-occurrence of its parent species. Morphologically it is rather variable but more similar
to B. rubrum, differing from the latter mainly in having leaves more or less farinose on the lower side.
Genus Kali Mill.
Kali tragus (L.) Scop. subsp. pontica (Pall.) Mosyakin, comb. nov.
Basionym: Salsola kali L. var. pontica Pall. 1803, Ill. Pl.: 37, tab. 29, fig. 2.
— Salsola kali L. subsp. pontica (Pall.) Mosyakin, 1996, Ann. Missouri Bot. Gard. 83(3): 389. —
S. tragus L. subsp. pontica (Pall.) S. Rilke, 1999, Biblioth. Bot. 149: 133. — S. pontica (Pall.) Degen,
1937, Fl. Velebit. 2: 47; Цвелев, 1993, Укр. ботан. журн. 50(1): 82.
In recent publications this taxon was recognized as a species (S. pontica), as a subspecies of S. kali,
or as a subspecies of S. tragus. My placement of this taxon in S. kali s. l. (Mosyakin, 1996, 2003) was
mostly caused by the assumption that S. tragus can be also treated eventually as a subspecies of S. kali.
However, S. pontica seems to be closer to or rooted in S. tragus (Rilke, 1999; Ryan et al., 2007).
Acknowledgments and dedication. I dedicate this article to the living memory of
my friend and co-author, Steven Earl Clemants (1954—2008), who greatly contrib-
uted to American and international plant conservation and taxonomic botany, espe-
cially taxonomy and geography of Chenopodiaceae, Amaranthaceae, Ericaceae,
Juncaceae and some other families.
REFERENCES
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diaceae): molecular phylogenetic analysis of nuclear and chloroplast data sets and a revised clas-
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4
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4
photosynthesis // Amer. J. Bot. — 2010. — 97. — P. 1664—1687.
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396 ISSN 0372-4123. Ukr. Botan. Journ., 2012, vol. 69, № 3
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Recommended for publication Submitted 15.06.2012 year
by D.V. Dubyna
С.Л. Мосякін
Інститут ботаніки імені М.Г. Холодного НАН України, м. Київ
НОВІ НОМЕНКЛАТУРНІ КОМБІНАЦІЇ В РОДАХ BLITUM L. І KALI MILL.
(CHENOPODIACEAE): ТАКСОНИ, ПРЕДСТАВЛЕНІ У ФЛОРІ СХІДНОЇ ЄВРОПИ
П’ять нових номенклатурних комбінацій (чотири на рівні секції й одна для гібрида) запро-
поновані в роді Blitum L. та одна нова комбінація на рівні підвиду — в роді Kali Mill. (Che no-
podiaceae). Стисло обґрунтована необхідність виділення цих родів зі складу Chenopodium L. і
Salsola L. відповідно.
К л ю ч о в і с л о в а: нові комбінації, Blitum, Kali, Chenopodium, Salsola.
С.Л. Мосякин
Институт ботаники имени Н.Г. Холодного НАН Украины, г. Киев
НОВЫЕ НОМЕНКЛАТУРНЫЕ КОМБИНАЦИИ В РОДАХ BLITUM L.
И KALI MILL. (CHENOPODIACEAE): ТАКСОНЫ, ПРЕДСТАВЛЕННЫЕ
ВО ФЛОРЕ ВОСТОЧНОЙ ЕВРОПЫ
Пять новых номенклатурных комбинаций (четыре на уровне секции и одна для гибрида)
предложены в роде Blitum L. и одна новая комбинация на подвидовом уровне — в роде Kali
Mill. (Chenopodiaceae). Кратко обоснована необходимость выделения этих родов из состава,
соответственно, Chenopodium L. и Salsola L.
К л ю ч е в ы е с л о в а: новые комбинации, Blitum, Kali, Chenopodium, Salsola.
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/ENU (Use these settings to create Adobe PDF documents best suited for high-quality prepress printing. Created PDF documents can be opened with Acrobat and Adobe Reader 5.0 and later.)
/UKR <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>
>>
/Namespace [
(Adobe)
(Common)
(1.0)
]
/OtherNamespaces [
<<
/AsReaderSpreads false
/CropImagesToFrames true
/ErrorControl /WarnAndContinue
/FlattenerIgnoreSpreadOverrides false
/IncludeGuidesGrids false
/IncludeNonPrinting false
/IncludeSlug false
/Namespace [
(Adobe)
(InDesign)
(4.0)
]
/OmitPlacedBitmaps false
/OmitPlacedEPS false
/OmitPlacedPDF false
/SimulateOverprint /Legacy
>>
<<
/AddBleedMarks false
/AddColorBars false
/AddCropMarks false
/AddPageInfo false
/AddRegMarks false
/ConvertColors /ConvertToCMYK
/DestinationProfileName ()
/DestinationProfileSelector /DocumentCMYK
/Downsample16BitImages true
/FlattenerPreset <<
/PresetSelector /MediumResolution
>>
/FormElements false
/GenerateStructure false
/IncludeBookmarks false
/IncludeHyperlinks false
/IncludeInteractive false
/IncludeLayers false
/IncludeProfiles false
/MultimediaHandling /UseObjectSettings
/Namespace [
(Adobe)
(CreativeSuite)
(2.0)
]
/PDFXOutputIntentProfileSelector /DocumentCMYK
/PreserveEditing true
/UntaggedCMYKHandling /LeaveUntagged
/UntaggedRGBHandling /UseDocumentProfile
/UseDocumentBleed false
>>
]
>> setdistillerparams
<<
/HWResolution [2400 2400]
/PageSize [612.000 792.000]
>> setpagedevice
|