Two New Spider Species of the Genus Draconarius (Araneae, Amaurobiidae, Coelotinae) from Pakistan
Draconarius pakistanicus Ovtchinnikov, sp. n. and D. naranensis Ovtchinnikov, sp. n. from Pakistan are described. The typical loci for Coelotes bifidus Caporiacco, 1935 and C. stylifer Caporiacco, 1935 are determined. Three species of the family Amaurobiidae (subfamily Coelotinae) are known now from...
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Інститут зоології ім. І. І. Шмальгаузена НАН України
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irk-123456789-33112009-09-01T17:34:03Z Two New Spider Species of the Genus Draconarius (Araneae, Amaurobiidae, Coelotinae) from Pakistan Ovtchinnikov, S.V. Inayatullah, M. Краткие сообщения Draconarius pakistanicus Ovtchinnikov, sp. n. and D. naranensis Ovtchinnikov, sp. n. from Pakistan are described. The typical loci for Coelotes bifidus Caporiacco, 1935 and C. stylifer Caporiacco, 1935 are determined. Three species of the family Amaurobiidae (subfamily Coelotinae) are known now from Pakistan: C. stylifer Caporiacco, 1935 was actually described from India, not from Pakistan. Из Пакистана описаны Draconarius pakistanicus Ovtchinnikov, 1999, sp. n. и D. naranensis Ovtchinnikov, 1999, sp. n. Для Coelotes bifidus Caporiacco, 1935 и C. stylifer Caporiacco, 1935 уточнены типовые локусы. Установлено, что в настоящее время в Пакистане известно 3 вида пауков из подсемейства Coelotinae (Amaurobiidae), а C. stylifer Caporiacco, 1935 описан не из Пакистана, а из Индии. 2005 Article Two New Spider Species of the Genus Draconarius (Araneae, Amaurobiidae, Coelotinae) from Pakistan / S. V. Ovtchinnikov, M. Inayatullah // Вестн. зоологии. — 2005. — Т. 39, № 3. — С. 85–88. — англ. 0084-5604 http://dspace.nbuv.gov.ua/handle/123456789/3311 595.44(549.1) en Інститут зоології ім. І. І. Шмальгаузена НАН України |
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Краткие сообщения Краткие сообщения Ovtchinnikov, S.V. Inayatullah, M. Two New Spider Species of the Genus Draconarius (Araneae, Amaurobiidae, Coelotinae) from Pakistan |
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Draconarius pakistanicus Ovtchinnikov, sp. n. and D. naranensis Ovtchinnikov, sp. n. from Pakistan are described. The typical loci for Coelotes bifidus Caporiacco, 1935 and C. stylifer Caporiacco, 1935 are determined. Three species of the family Amaurobiidae (subfamily Coelotinae) are known now from Pakistan: C. stylifer Caporiacco, 1935 was actually described from India, not from Pakistan. |
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Ovtchinnikov, S.V. Inayatullah, M. |
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Ovtchinnikov, S.V. Inayatullah, M. |
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Ovtchinnikov, S.V. |
| title |
Two New Spider Species of the Genus Draconarius (Araneae, Amaurobiidae, Coelotinae) from Pakistan |
| title_short |
Two New Spider Species of the Genus Draconarius (Araneae, Amaurobiidae, Coelotinae) from Pakistan |
| title_full |
Two New Spider Species of the Genus Draconarius (Araneae, Amaurobiidae, Coelotinae) from Pakistan |
| title_fullStr |
Two New Spider Species of the Genus Draconarius (Araneae, Amaurobiidae, Coelotinae) from Pakistan |
| title_full_unstemmed |
Two New Spider Species of the Genus Draconarius (Araneae, Amaurobiidae, Coelotinae) from Pakistan |
| title_sort |
two new spider species of the genus draconarius (araneae, amaurobiidae, coelotinae) from pakistan |
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Інститут зоології ім. І. І. Шмальгаузена НАН України |
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2005 |
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Краткие сообщения |
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Two New Spider Species of the Genus Draconarius (Araneae, Amaurobiidae, Coelotinae) from Pakistan / S. V. Ovtchinnikov, M. Inayatullah // Вестн. зоологии. — 2005. — Т. 39, № 3. — С. 85–88. — англ. |
| work_keys_str_mv |
AT ovtchinnikovsv twonewspiderspeciesofthegenusdraconariusaraneaeamaurobiidaecoelotinaefrompakistan AT inayatullahm twonewspiderspeciesofthegenusdraconariusaraneaeamaurobiidaecoelotinaefrompakistan |
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2025-07-02T06:37:57Z |
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2025-07-02T06:37:57Z |
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UDC 595.44(549.1)
TWO NEW SPIDER SPECIES OF THE GENUS DRACONARIUS
(ARANEAE, AMAUROBIIDAE, COELOTINAE) FROM PAKISTAN
S. V. Ovtchinnikov1, M. Inayatullah2
1 Scientific group “Entomologist”, Consultation and Information Society “Biosphere”,
Maldybaeva, 9/32, Bishkek, 720055 Kyrgyzstan
2 Entomological Department, Agricultural University, NWFP, Peshawar, Pakistan
Accepted 15 June 2004
Two New Spider Species of the Genus Draconarius (Araneae, Amaurobiidae, Coelotinae) from Pakistan.
Ovtchinnikov S. V., Inayatullah M. — Draconarius pakistanicus Ovtchinnikov, sp. n. and D. naranensis
Ovtchinnikov, sp. n. from Pakistan are described. The typical loci for Coelotes bifidus Caporiacco, 1935
and C. stylifer Caporiacco, 1935 are determined. Three species of the family Amaurobiidae (subfamily
Coelotinae) are known now from Pakistan: C. stylifer Caporiacco, 1935 was actually described from
India, not from Pakistan.
Ke y wo r d s: Araneae, Amaurobiidae, Draconarius, Coelotes, new species, Pakistan.
Äâà íîâûõ âèäà ðîäà Draconarius (Araneae, Amaurobiidae, Coelotinae) èç Ïàêèñòàíà. Îâ÷èííèêîâ Ñ. Â.,
Èíàÿòóëëà Ì. — Èç Ïàêèñòàíà îïèñàíû Draconarius pakistanicus Ovtchinnikov, 1999, sp. n. è
D. naranensis Ovtchinnikov, 1999, sp. n. Äëÿ Coelotes bifidus Caporiacco, 1935 è C. stylifer Caporiacco,
1935 óòî÷íåíû òèïîâûå ëîêóñû. Óñòàíîâëåíî, ÷òî â íàñòîÿùåå âðåìÿ â Ïàêèñòàíå èçâåñòíî 3
âèäà ïàóêîâ èç ïîäñåìåéñòâà Coelotinae (Amaurobiidae), à C. stylifer Caporiacco, 1935 îïèñàí íå
èç Ïàêèñòàíà, à èç Èíäèè.
Êëþ÷åâûå ñ ëîâ à: Araneae, Amaurobiidae, Draconarius, Coelotes, íîâûå âèäû, Ïàêèñòàí.
Up to the present time only one species of the spider subfamily Coelotinae (Ama-
urobiidae) has been reported from Pakistan — Paracoelotes bidens (Caporiacco, 1935),
described from Karakorum (Baltistan, near Baltoro Glacier, 3,900–4,150 m. a. s. l.,
35°41'N, 76°09'E) and NW Himalaya (Tolti environs, 2,400 m. a. s. l., 35°02'N,
76°01'E). Type locality of the second coelotin species (Coelotes stylifer Caporiacco,
1935) described in the same paper is India (Jammu and Kashmir State, Gund environs,
2,080 m. a. s. l., 34°16'N, 75°05'E, and Kangan, 2,000 m. a. s. l., 34°15'N, 74°55'E).
Two new species of the genus Draconarius Ovtchinnikov, 1999 were found by us
in 2003 in W Himalaya, both of which are clearly closed to the type species of the
genus, D. venustus Ovtchinnikov, 1999 from Gissaro-Darvaz Mountain system, from
where we have another, undescribed species.
In the present article these two new species from Western Himalaya are described.
The following abbreviations have been used in the text: ALE — anterior lateral eyes;
AME — anterior median eyes; PLE — posterior lateral eyes; PME — posterior medi-
an eyes; PLS — posterior lateral spinnerets. All measurements are in millimetres. The
scale unit on all figures equals 0.5 mm. All holotypes and part of paratypes will be
deposited in the Zoological Museum of Moscow State University, Moscow (Dr. K. G. Mi-
khailov); remaining paratypes are in the private collection of the first author (Bishkek,
Kyrgyzstan).
Draconarius pakistanicus Ovtchinnikov, sp. n. (fig. 1–4)
Type ma t e r i a l. Holotype {, Pakistan, W Himalaya, 10 km SE from Babusar Pass, north slope,
4000 m a. s. l., 10.08.2003. Paratypes: 14 {, 16 }, same data as holotype (Ovtchinnikov).
Vestnik zoologii, 39(3): 85–88, 2005
© S. V. Ovtchinnikov, M. Inayatullah, 2005
86 S. V. Ovtchinnikov, M. Inayatullah
Fig. 1–4. Draconarius pakistanicus: 1 — left palpus, ventral view; 2 — ditto, retrolateral view; 3 — epigynum,
ventral view; 4 — ditto, dorsal view.
Ðèñ. 1–4. Draconarius pakistanicus: 1 — ëåâûé ïàëüïóñ, âåíòðàëüíî; 2 — òî æå, ðåòðîëàòåðàëüíî; 3 —
ýïèãèíà, âåíòðàëüíî; 4 — òî æå, äîðñàëüíî.
Dia gno s i s. Copulative structures in both sexes similar to those in D. venustus
Ovtchinnikov. Male differs by great size, relatively small cymbium of the palpus and
correspondingly less developed apophysises of the conductor, and not so long embolus
(fig. 1, 2). Females of both species have externally similar epigynae and spermathecae,
as well as proportions and the angle of relative position (fig. 3, 4). The new species has
fewer convolutions of the copulatory ducts, therefore the last are wider. Spiders of the
new species are about twice bigger, and have contrastly bicoloured legs I–III. The new
species differs from the geographically closest species D. naranensis sp. n. by absence of
transverse stripes on legs, by absence of maculae on carapace and sternite, and by shape
and relative position of epigynal teeth. The new species is close by male palpus also to
D. aspinatus (Wang & al., 1990) from Anhui Province, China. But the last one has
shorter cymbial apex and conductor, which not twisted on its end. Besides, it has not
patellar apophysis.
De s c r i p t i o n. Male: Total body length 14 (holotype), 11–14 (paratypes). Length
of carapace 6.30, width of carapace 4.10, with ratio 1.55; length of head part 1.7, its
width in area of PER 0.64 times maximal width of thorax. Relative diameters of an eye
and distance between them (in standard units) AME 10, ALE 16, PME 12, PLE 15,
AME–AME 10, ALE–AME 9, ALE-PLE 7, PLE–PME 19, PME–PME 14. Sternum
length 3.1, width 2.25. Hind tibia + patella to length of carapace 1.11. Leg formula
oftenest 4123, commonly 4321, sometimes 4132. Carapace yellow-brown, slightly dark-
ened in the anterior part of head. Metatarsi I-III entirely black-brown; tibiae I and II
darkened only partially. Other parts of legs yellow-brown except fore femora and coxae
which are somewhat darker. Carapace, sternum and legs devoid of pigment macula-
tions. Carapace clothed in black adpressed hairs with obliquely truncated apex; similar
hairs on femora, patellae, tibiae, and bases of metatarsi. Abdomen and sternum in sim-
ple hairs. Pattern on abdomen grey, not sharply delineated, prevailing against the white
background. Spinnerets of moderate size, apical to basal PLS segment ratio 0.8.
F ema l e. Body length 11.5–17. Carapace length 4.5, width 3. Width of head to
that of carapace 0.7. Apical segment of PLS to its basal segment 0.95. Epigynum of
common type (fig. 3–4). Epigynal teeth are located above fovea, not extending outside
its margins. Sperm receptacle ducts form oblong oval coils, pointing towards to each
other and adjoin by the apices. Ducts wide, of light colour, delimited below by dark
stripes. Spermathecae twice shorter than ducts. Their inner margins delimited by nar-
row, white bursae copulatrices (tabl. 1).
E co l o g y. D. pakistanicus sp. n. abundantly occurs in subalpine and alpine mead-
ows, its biomass prevailing over most of other spiders. Provided sufficient humidity, it
can be found easily under any boulder. It is one of the biggest-sized species, and the
sole representative of its family in the area of the research.
D i s t r i b u t i o n. Pakistan: Western Himalaya.
E t ymo l o g y. Named after the country name in which it was collected.
Draconarius naranensis Ovtchinnikov, sp. n. (fig. 5, 6)
Typ e ma t e r i a l. Holotype $, Pakistan, W Himalaya, ca. 3 km NNE Naran, 2,500 m a. s. l.,
coniferous forest (Cedrus, Picea, Pinus) 18.08.2003. Paratype: $, same data as holotype (S. V. Ovtchinnikov).
87Two New Spider Species of the Genus Draconarius…
Ta b l e 1. Length of legs (male/female)
Ò à á ëèö à 1. Äëèíà ÷ëåíèêîâ íîã (ñàìåö/ñàìêà)
I 5.25/4.50 2.25/2.25 4.90/4.15 5.60/4.2 3.35/2.25 21.12/17.35
II 5.10/4.40 2.10/2.25 4.35/3.60 5.45/4.2 3.10/2.10 20.10/15.55
III 4.85/3.85 2.10/2.20 3.75/3.15 5.45/4.40 2.85/2.15 19.00/15.75
IV 5.60/4.95 2.35/2.30 5.05/4.45 7.25/5.95 3.35/2.65 23.60/20.25
Legs Femur Patella Tibia Metatarsus Tarsus Total
Dia gno s i s. Very close to D. pakistanicus sp. n. by the structure and position of
copulatory ducts and spermathecae, but differs from it by the external structure of epig-
ynum. (fig. 5,6). Epigynal teeth of D. naranensis sp. n., in difference from the previous
species, are weakly sclerotized, poorly pigmented and longer. It also is smaller and more
lightly coloured, has all legs uniformly testaceous and femora transversely striped.
De s c r i p t i o n. Total body length 8.4 (holotype), 8.8 (paratype). Length of cara-
pace 4.0–4.2, width of carapace 2.83–2.88, length: width ratio 1.39–1.48. Length of
head part 0.98–1.00, width 1.75–1.88, width of head part to width of carapace
0.61–0.66. Relative diameters of an eye and distance between them (in standard units)
AME 12, PME 20, PLE 14; AME–AME 8, AME–ALE 6, ALE–PLE 5, PLE-PME
12, PME-PME 11. Length of carapace to length of patella + tibia 1.07–1.16 (tabl. 2).
Ca r a p a c e yellow-brown with darkened anterior part and black-grey marginal line
around thoracic region. Radiating from fissure of carapace are 4 elongated spots, pairs
2–4 of which broadening peripherally towards sides where they are delimited by a light
marginal stripe encircling thoracic region. Legs yellow-brown with weak bands of lighter
colour. Abdomen with a typical amaurobiid pattern.
D i s t r i b u t i o n. Pakistan: Western Himalaya.
E t ymo l o g y. Toponymic, after its type locality.
Caporiacco L. di. Aracnidi dell’ Himalaia e del Karakoram, raccolti dalla Missione italiana al Karakoram
(1929-VII) // Mem. Soc. ent. ital. — 1935. — 13. — P. 161–263.
Ovtchinnikov S. V. On the supraspecific systematics of the subfamily Coelotinae (Araneae, Amaurobiidae) in
the former USSR fauna // Tethys entomol. Research. — 1999. — 1. — P. 63–79.
88 S. V. Ovtchinnikov, M. Inayatullah
Fig. 5–6. Draconarius naranensis: 5 — epigynum, ventral view; 6 — ditto, dorsal view.
Ðèñ. 5–6. Draconarius naranensis: 5 — ýïèãèíà, âåíòðàëüíî; 6 — òî æå, äîðñàëüíî.
Ta b l e 2. Length of legs (holotype/paratype)
Ò à á ëèö à 2. Äëèíà ÷ëåíèêîâ íîã (ãîëîòï/ïàðàòèï)
I 3.35/3.33 1.53/1.43 3.10/3.05 3.18/3.15 1.75/1.73 12.91/12.69
II 3.20/3.05 1.43/1.33 2.65/2.65 2.88/2.85 1.58/1.58 11.74/11.46
III 2.88/2.78 1.25/1.20 2.20/2.18 2.83/2.80 1.38/1.40 10.54/10.36
IV 3.60/3.58 1.38/1.35 3.15/3.10 3.75/3.85 1.63/1.70 13.51/13.58
Legs Femur Patella Tibia Metatarsus Tarsus Total
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