New and Little-Known Tephritidae (Diptera, Cyclorrhapha) from Europe
Описаны два новых вида трибы Terelliini (Tephritinae). Terellia (Cerajocera) cyanoides Korneyev sp. n. с северо-востока Украины отличается от других видов подрода комбинацией полосатого рисунка крыла, невидоизмененного педицеллюма, катепистернума с черным пятном и малыми размерами тела; личинки бура...
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Інститут зоології ім. І. І. Шмальгаузена НАН України
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irk-123456789-37812009-09-01T17:31:03Z New and Little-Known Tephritidae (Diptera, Cyclorrhapha) from Europe Korneyev, V.A. Фауна и систематика Описаны два новых вида трибы Terelliini (Tephritinae). Terellia (Cerajocera) cyanoides Korneyev sp. n. с северо-востока Украины отличается от других видов подрода комбинацией полосатого рисунка крыла, невидоизмененного педицеллюма, катепистернума с черным пятном и малыми размерами тела; личинки буравят стебли Jurinea sp. ex cyanoides aggr. на песчаных холмах в сосновом бору. Orellia tragopogonis Korneyev et Dirlbek sp. n. из Испании близка O. scorzonerae (Robineau-Desvoidy), отличаясь соединенными апикальной и преапикальной перевязями крыла, передним бедром, покрытым дорсально белыми волосками, а также большими размерами; личинки питаются в соцветиях Tragopogon dubium. 14 видов Tephritidae впервые отмечены из ряда стран Европы. Приведены таблицы для определения видов Terellia подрода Cerajocera и рода Orellia. Two new species of the tribe Terelliini (Tephritinae) are described. Terellia (Cerajocera) cyanoides Korneyev sp. n. from northeastern Ukraine differs from other species of the subgenus by combination of banded wing, unmodified pedicel, black-spotted katepisternum and small size; its larvae bore stems of Jurinea sp. ex cyanoides aggr. on sand dunes in pine forests. Orellia tragopogonis Korneyev et Dirlbek sp. n. from Spain fits near O. scorzonerae (Robineau-Desvoidy), differing by fused apical and preapical wing crossbands, fore femur dorsally white setose and by larger size; its larvae feed in flower heads of Tragopogon dubium. 14 species of Tephritidae are recorded for the first time from certain European countries. Keys to species of Terellia subgenus Cerajocera and Orellia are provided. 2003 Article New and Little-Known Tephritidae (Diptera, Cyclorrhapha) from Europe / V. A. Korneyev // Вестн. зоологии. — 2003. — Т. 37, № 3. — С. 3-12. — англ. 0084-5604 http://dspace.nbuv.gov.ua/handle/123456789/3781 595.773.4 en Інститут зоології ім. І. І. Шмальгаузена НАН України |
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Фауна и систематика Фауна и систематика Korneyev, V.A. New and Little-Known Tephritidae (Diptera, Cyclorrhapha) from Europe |
| description |
Описаны два новых вида трибы Terelliini (Tephritinae). Terellia (Cerajocera) cyanoides Korneyev sp. n. с северо-востока Украины отличается от других видов подрода комбинацией полосатого рисунка крыла, невидоизмененного педицеллюма, катепистернума с черным пятном и малыми размерами тела; личинки буравят стебли Jurinea sp. ex cyanoides aggr. на песчаных холмах в сосновом бору. Orellia tragopogonis Korneyev et Dirlbek sp. n. из Испании близка O. scorzonerae (Robineau-Desvoidy), отличаясь соединенными апикальной и преапикальной перевязями крыла, передним бедром, покрытым дорсально белыми волосками, а также большими размерами; личинки питаются в соцветиях Tragopogon dubium. 14 видов Tephritidae впервые отмечены из ряда стран Европы. Приведены таблицы для определения видов Terellia подрода Cerajocera и рода Orellia. |
| format |
Article |
| author |
Korneyev, V.A. |
| author_facet |
Korneyev, V.A. |
| author_sort |
Korneyev, V.A. |
| title |
New and Little-Known Tephritidae (Diptera, Cyclorrhapha) from Europe |
| title_short |
New and Little-Known Tephritidae (Diptera, Cyclorrhapha) from Europe |
| title_full |
New and Little-Known Tephritidae (Diptera, Cyclorrhapha) from Europe |
| title_fullStr |
New and Little-Known Tephritidae (Diptera, Cyclorrhapha) from Europe |
| title_full_unstemmed |
New and Little-Known Tephritidae (Diptera, Cyclorrhapha) from Europe |
| title_sort |
new and little-known tephritidae (diptera, cyclorrhapha) from europe |
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Інститут зоології ім. І. І. Шмальгаузена НАН України |
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2003 |
| topic_facet |
Фауна и систематика |
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http://dspace.nbuv.gov.ua/handle/123456789/3781 |
| citation_txt |
New and Little-Known Tephritidae (Diptera, Cyclorrhapha) from Europe / V. A. Korneyev // Вестн. зоологии. — 2003. — Т. 37, № 3. — С. 3-12. — англ. |
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2025-07-02T07:01:33Z |
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1836517628051456000 |
| fulltext |
UDC 595.773.4
NEW AND LITTLE-KNOWN TEPHRITIDAE
(DIPTERA, CYCLORRHAPHA) FROM EUROPE
V. A. Korneyev
Schmalhausen Institute of Zoology, National Academy of Sciences of the Ukraine,
Vul. B. Khmelnitskogo, 15, Kyiv, MSP, 01601 Ukraine
E-mail: korval@entom.freenet.kiev.ua
Accepted 5 December 2002
New and Little-Known Tephritidae (Diptera, Cyclorrhapha) from Europe. Korneyev V. A. — Two new
species of the tribe Terelliini (Tephritinae) are described. Terellia (Cerajocera) cyanoides Korneyev sp. n.
from northeastern Ukraine differs from other species of the subgenus by combination of banded wing,
unmodified pedicel, black-spotted katepisternum and small size; its larvae bore stems of Jurinea sp. ex
cyanoides aggr. on sand dunes in pine forests. Orellia tragopogonis Korneyev et Dirlbek sp. n. from Spain
fits near O. scorzonerae (Robineau-Desvoidy), differing by fused apical and preapical wing crossbands,
fore femur dorsally white setose and by larger size; its larvae feed in flower heads of Tragopogon dubium.
14 species of Tephritidae are recorded for the first time from certain European countries. Keys to
species of Terellia subgenus Cerajocera and Orellia are provided.
Ke y wo r d s: Diptera, Tephritidae, Europe, new species, new records.
Íîâûå è ìàëîèçâåñòíûå Tephritidae (Diptera, Cyclorrhapha) èç Åâðîïû. Êîðíååâ Â. À. — Îïèñàíû
äâà íîâûõ âèäà òðèáû Terelliini (Tephritinae). Terellia (Cerajocera) cyanoides Korneyev sp. n. ñ ñå-
âåðî-âîñòîêà Óêðàèíû îòëè÷àåòñÿ îò äðóãèõ âèäîâ ïîäðîäà êîìáèíàöèåé ïîëîñàòîãî ðèñóíêà
êðûëà, íåâèäîèçìåíåííîãî ïåäèöåëëþìà, êàòåïèñòåðíóìà ñ ÷åðíûì ïÿòíîì è ìàëûìè ðàçìå-
ðàìè òåëà; ëè÷èíêè áóðàâÿò ñòåáëè Jurinea sp. ex cyanoides aggr. íà ïåñ÷àíûõ õîëìàõ â ñîñíîâîì
áîðó. Orellia tragopogonis Korneyev et Dirlbek sp. n. èç Èñïàíèè áëèçêà O. scorzonerae (Robineau-
Desvoidy), îòëè÷àÿñü ñîåäèíåííûìè àïèêàëüíîé è ïðåàïèêàëüíîé ïåðåâÿçÿìè êðûëà, ïåðåäíèì
áåäðîì, ïîêðûòûì äîðñàëüíî áåëûìè âîëîñêàìè, à òàêæå áîëüøèìè ðàçìåðàìè; ëè÷èíêè ïè-
òàþòñÿ â ñîöâåòèÿõ Tragopogon dubium. 14 âèäîâ Tephritidae âïåðâûå îòìå÷åíû èç ðÿäà ñòðàí Åâ-
ðîïû. Ïðèâåäåíû òàáëèöû äëÿ îïðåäåëåíèÿ âèäîâ Terellia ïîäðîäà Cerajocera è ðîäà Orellia.
Êëþ÷åâûå ñ ëîâ à: Diptera, Tephritidae, Åâðîïà, íîâûå âèäû, íîâûå íàõîäêè.
Introduction
The fruit flies (Tephritidae) are one of the most economically important Diptera families, first of all,
in tropics and subtropics. In Palaearctic Region, and especially in Europe, this family is represented by the
asparagus fly Plioreocepta poeciloptera (Schrank) and a few dozens of fruit-feeding species, including the
medfly Ceratitis capitata (Wiedemann), the cherry fruit fly Rhagoletis cerasi (Linnaeus), but the vast majority
of Palearctic species, which belong in the subfamily Tephritinae, feed in flowerheads and stems (often
forming galls) on plants of the family Asteraceae. European fauna is comparatively well known, except for
Balkan countries and some regions of Eastern Europe.
Dr. Bernhard Merz (Museum d'Histoire Naturelle, Geneve) kindly sent me his manuscript of the
Tephritidae chapter of the Fauna Europaea Dipteran volume (Merz, in press) for preliminary reviewing.
Comparison of the distributional data against the material deposited in my collection (CVK) and in
collections of the I. I. Schmalhausen Institute of Zoology (Kyiv), Deutsche Entomologisches Institut,
Eberswalde (DEI), Termeszettudományi Muzeum, Budapest (TMB), Naturhistorisches Museum Wien
(NHMW), Narodni Muzeum v Praze (NMP), Zoological Museum of Russian Academy of Sciences, St.
Petersburg (ZISP) and Zoological Museum of the M. V. Lomonosov University, Moscow (ZMUM), added
a few new data based upon the material listed below.
Morphological terminology follows J. F. McAlpine (1981).
Euphranta connexa (Fabricius, 1794)
Ma t e r i a l e x am in ed. Moldova, Goian, 14.06.1986, 2 {, 4 } (Korneyev) (CVK).
Commen t s. First record from Moldova.
Vestnik zoologii, 37(3): 3–12, 2003
© V. A. Korneyev, 2003 Ôàóíà è ñèñòåìàòèêà
Trypeta zoe Meigen, 1826
Ma t e r i a l e x am in ed. Belarus: vicinity of Minsk, 23.05.1979, 2 { (V. Nazarov) (CVK).
Commen t s. First record from Belarus.
Rhagoletis cerasi (Linnaeus, 1758)
Ma t e r i a l e x am in ed. Croatia: "Split. D[almatien]" 21.04.1961, } (TMB). Greece, Makedonia,
Kilkis, Koupa, 5.06.2002, 2 {, 3 } (Korneyev) (CVK).
Commen t s. First record from Croatia and Greece.
Terellia (Cerajocera) euura (Hering) (fig. 1)
Ma t e r i a l e x am in ed. Holotype }, SE of European, Russia, "Had", "293", "coll. // H. Loew",
"Sarepta // Christoph" [latest cyan label], "eu- // ura // Lw. " [Loew's writing], "Type" [red paper label added
by Enderlein] (ZMHB).
Commen t s. This species is known only from its type locality. Further records
from Ukraine and Uzbekistan (Richter, 1970) are based upon misidentified specimens
of Chaetostomella cylindrica Robineau-Desvoidy ("Aleshki Kherson [Gov.],
12.07.1924", {) (ZISP), and Terellia (Cerajocera) sp. near maculicauda Chen ("Katta-
Kurgan, 16.05.1929") (ZISP).
4 V. A. Korneyev
Fig. 1. Terellia (Cerajocera) euura (Loew), holotype: 1 — total view, dorsally; 2 — aculeus tip, ventral view,
enlarged; 3 — head, lateral view; 4 — face (antennae not shown).
Ðèñ. 1. Terellia (Cerajocera) euura (Loew), ãîëîòèï: 1 — îáùèé âèä, äîðñàëüíî; 2 — âåðøèíà àêóëåóñà,
âåíòðàëüíî, óâåëè÷åíî; 3 — ãîëîâà, ñáîêó; 4 — ëèöî (àíòåííû íå ïîêàçàíû).
Fig. 2. Terellia (Cerajocera) cyanoides: 1 — head, lateral view; 2 — wing; 3 — male abdomen; 4 — glans of
phallus; 5 — aculeus; 6 — apex, enlarged; 7 — puparium in stem; 8 — caudal segment of puparium. Scale
bar 1 mm.
Ðèñ. 2. Terellia (Cerajocera) cyanoides: 1 — ãîëîâà, âèä ñáîêó; 2 — êðûëî; 3 — áðþøêî ñàìöà; 4 — ãëàíñ
ôàëëþñà; 5 — àêóëåóñ; 6 — âåðøèíà, óâåëè÷åíî; 7 — ïóïàðèé; 8 — êàóäàëüíûé ñåãìåíò ïóïàðèÿ.
Ìàñøòàáíàÿ ëèíåéêà 1 ìì.
Terellia (Cerajocera) tussilaginis (Fabricius)
Ma t e r i a l e x am in ed. Estonia, Koeru; Peedu, 2 {, 2 } (Stackelberg); Lithuania, Kaunas ("Kow-
no"), } (ZISP).
Commen t s. First record from Estonia and Lithuania.
Terellia (Cerajocera) cyanoides Korneyev, sp. n. (fig. 2)
Ma t e r i a l e x am in ed. Holotype {, Ukraine, Chernihivs'ka Oblast', Yaduty 40 km NW of Bakhmach,
sand dunes in a pine forest, from stems of Jurinea sp. ex cyanoides aggr., 17.08.2000 — exit 1.03.2001
(V.Korneyev) (SIZK). Paratypes: {, 3 } (aculeus exposed in 2 }), same label as in the holotype (CVK,
SIZK).
De s c r i p t i o n. Body yellow to brownish-yellow in dry specimens, somewhat gre-
enish-yellow in live specimens.
Ma l e. Head (fig. 2, 1) yellow, except for ocellar triangle black. Head length :
height : width ratio = 1 : 1.13 : 1.43. Scapus and pedicel uniformly yellow, without
darker marks, scapus with 6–8 blackish setulae at dorso-apical margin, pedicel with thin
5New and Little-Known Tephritidae from Europe
and short black setulae: 4–7 on dorso-lateral, 12–15 on dorso-medial surface (1 longer
seta plus 3–4 irregular rows of shorter setulae) and 9–12 setulae forming single row at
apico-ventral margin. Flagellomere 1 light yellow, 1.8 times longer than wide. Arista
bare, 1.6–1.7 times longer than flagellomere 1, yellow in basal 0.25 of its length, dark
brown in the rest. Distance between bases of antennae 0.3 of scapus width. Palpus not
extending beyond anterior oral margin, pale yellow, with white setulae in basal 0.8 and
6–8 black setulae at apex. Proboscis capitate. with rather fine pale brown and whitish
setulae. Frons as long as wide at posterior margin, narrowed towards anterior margin;
frontal plates with 4–7 black setulae at posterior 0.2 of its length, and white setulose in
remaining portion. Parafacialium at narrowest place 0.3 as wide as flagellomere 1.
Facialium and gena with 2 rows of blackish setulae. Postgena with white setae below
and posterior of black and long genal seta. Eye vertical, 1.21 times higher than long.
Gena narrow, 0.14 as long as eye vertical diameter. Head setae dark brown to black,
except for postocellar, postvertical, postocular and occipital setae all white.
Thorax densely white setulose, with setae dark brown to black. Mesonotum dull,
short, but densely microtrichose, with common black lyrate pattern. Medial vitta
exceeding dorsocentral setae posteriorly. Both dorsocentral and acrostichal seta with
black alveoli and black spots around them. Scutellum yellow, flattened. with
12–16white setulae at each side and 4 setae; antero-ventral corners at squamal ridge
blackish. Pleura subshining yellow; katepisternum with large triangular dull black spot;
meron at middle, katatergite and anatergite at posteroventral margin dull black.
Postnotum dull black.
Wing (fig. 2, 2) with 4 narrow brownish-grey crossbands. Vein R4+5 with 1–3 setu-
lae basally on dorsal surface and 0–1 on ventral surface. Subbasal crossband restricted
to grey spot anterior of radial fork in r1 cell and in br cell anterior of BM–Cu crossvein;
costal cell hyaline or with inconspicuous shadow of grey. Cell sc yellow, except the very
apex grey; discal and preapical crossbands narrow, at most as wide as R–M vein length;
apical crossband separated from discal, in cell r4+5 narrower than distance between
apices of R4+5 and M vein. Postero-apical lobe of bcu not exceeding BM–Cu vein level.
Legs yellow, fore femur thickened in male, with 7–8 white setae in dorso-basal and
black setae in remaining portion.
Abdominal tergite 1+2 white setulose with a few black latero-marginal setae, ter-
gites 3 and 4 white setulose with single row of black marginal setae each, tergite 5 with
numerous black setae in medially in posterior half of its length. Tergites 3–5 with
4 black spots at anterior margin each, and 2 additional postero-lateral spots on tergite
5 separated from antero-lateral spots (fig. 2, 3).
Epandrium not dissected for study. Phallus glans (fig. 2, 4) like in other species of
Terellia (Cerajocera).
Female similar to male but fore femur non-thickened. Tergites 3–5 white setulose
with single row of black marginal setae, tergite 6 with 7–8 long marginal setae, black
setulose in posterior 1/3 in medial portion, 1.5 times as long as tergite 5. Tergosternite
7 yellow with 2 large dark brown spots basally, black setulose, slightly longer than ter-
gites 3–6 together. Aculeus rounded apically (fig. 2, 5, 6), twice as long as costal cell.
Measurements (length in mm). Male. Body 3.9–4.2, wing 3.5–3.7. Female. Body
(with oviscape) 4.6–5.6, wing 3.5–4.0, oviscape dorsally 1.4–1.5, aculeus 1.7–1.8.
Puparium. Yellow with black caudal segment and one simple sclerotized prong
below posterior spiracles at middle (fig. 2, 8).
B i o l o g y. Larvae feed in flower heads of Jurinea, and later bore stems and pupate
inside them in cocoons made of plant remainders (fig. 2, 7).
The new species can be recognized from other species of the subgenus Cerajocera
by the following key.
6 V. A. Korneyev
Key to species of Terellia (Cerajocera)
Òàáëèöà äëÿ îïðåäåëåíèÿ âèäîâ Terellia (Cerajocera)
1. Wings hyaline. Caudal segment of 3rd instar larva and puparium with single sclerotized process ...
....................................................................................................................................................... 2.
– Wings dark banded or spotted. Caudal segment various (with single, double sclerotized process or
without it) .................................................................................................................................... 3.
2. Tergites 3 and 4 with black setulae both medially in their discs and along posterior margins. Larvae
in Jurinea mollis aggr. stems ..........................................................................T. (C.) setifera Hendel
– Tergites 3 and 4 yellow setulose except for black marginal setulae on tergite 4 and a few latero-
marginal setulae on tergite 3. Larvae in Jurinea cyanoides aggr. stems ............................................
.............................................................................................................. T. (C.) clarissima Korneyev
3. Distance between bases of antennae exceeding diameter of scapus; medial lobe of pedicel enlarged
in females or with large spinulose lobe in males. Tergite 4 usually black setulose. Caudal segment
of larva with 2 separate processes or one fish-tail-like process (unknown for T. (C.) armeniaca) ..
....................................................................................................................................................... 4.
– Antennae closer together than diameter of scapus. Tergite 4 white setulose. Caudal segment with
simple process or without it. ........................................................................................................ 6.
4. Subbasal crossband well-developed from costal to anal vein; costal cell distinctly infuscate. Larvae
in flower heads of Centaurea scabiosa aggr. Process of caudal segment of larva with fish-tail apex.
................................................................................................................. T. (C.) ceratocera Hendel
– Subbasal crossband poorly developed or absent; costal cell completely hyaline .......................... 6.
5. Discal and preapical crossbands exceeding medial vein and usually reaching cubital vein. Larvae in
stems of Centaurea scabiosa aggr. Caudal segment of larva with 2 separate processes ....................
.............................................................................................................. T. (C.) plagiata (Dahlbom)
– Discal crossband absent, preapical crossband posteriorly reaching at most to medial vein. Larvae
unknown ............................................................................................. T. (C.) armeniaca Korneyev
6. Katepisternum and meron with reddish-brown spots. Larvae in seeds of Arctium spp. Caudal
segment of larva without sclerotized processes ............................................................................. 7.
– Katepisternum and meron with black spots. Larvae in receptacles or stems. Caudal segment of larva
usually with single sclerotized process (fig. 2, 8) ......................................................................... 8.
7. Subcostal cell yellow, brown at most in apical half; posterior portion of discal and anterior portion
of preapical crossbands distinct, yellow, brown bordered. Katepisternum with reddish, rarely black,
spot. Mesonotal pattern in anterior half usually reddish to brown ...................................................
.......................................................................................................... T. (C.) tussilaginis (Fabricius)
– Subcostal cell all brown; posterior portion of discal and anterior portion of preapical crossbands,
pale yellow, indistinctly bordered. Katepisternum with black spot. Mesonotal pattern black. .........
........................................................................................................... T. (C.) nigronota (Korneyev)
8. Subbasal crossband well-developed from costal to anal vein; costal cell widely infuscate or yellow
....................................................................................................................................................... 9.
– Subbasal crossband poorly developed or absent; costal cell completely hyaline or at most with faint
shadow of grey ............................................................................................................................ 10.
9. Tergite 6 of female white setulose. Major setae usually yellow to light brown in Eastern Europe or
black in Western Europe and Near East. Larvae in stems (Eastern and Central Europe) or in
receptacles (Western Europe) of Onopordum spp .............................................................................
.................................. T. (C.) gynaecochroma (Hering) (= T. (C.) lappae auctt., nec Cederhjelm)
– Tergite 6 of female black setulose, marginal setae black (fig. 1, 1). Larvae and host plants unknown.
South eastern Europe (Russia: Volgograd) ................................................. T. (C.) euura (Hering)
10. Discal and preapical crossbands wide, in cell r2+3 as wide as hyaline space between them. Larvae
in receptacles of Rhaponticum scariosum (Swiss and Italian Alps) ............ T. (C.) rhapontici Merz
– Discal and preapical crossbands narrow, in cell r2+3 at most 0.7 as wide as hyaline space between
them (fig. 2, 2). Larvae in stems ................................................................................................ 11.
11. Wing length more than 4.3 mm (4.6–5.4). Larvae in stems of Saussurea spp. in Kyrghyzia
(Korneyev, unpublished data) ............................................................ T. (C.) maculicauda (Chen)
– Wing length less than 4.0 mm (3.5–3.7). Larvae in stems of Jurinea cyanoides aggr. (Ukraine) ...
................................................................................................... T. (C.) cyanoides Korneyev, sp. n.
7New and Little-Known Tephritidae from Europe
Orellia tragopogonis Korneyev et J. Dirlbek, sp. n. (fig. 3)
Ma t e r i a l e x am in ed. Holotype }, "Hispania, Benidorm, 19.7–2.8. 1982, leg. J. Dirlbek", "kvety
[flowers of] Tragopogon dubius" [sic] (aculeus exposed) (NMP). Paratype }, "Spanien, Granada, 27.04.1974,
lgt. Dirlbek J. " (abdomen dissected) (CVK). 5 flower heads of Tragopogon dubium with cocoons and pupa-
ria, "Spanien, Granada, 27.04.1974, lgt. Dirlbek J." (CVK; NMP)
De s c r i p t i o n. Body yellow to brownish-yellow.
F ema l e. Head (fig. 3, 1) yellow, except for ocellar triangle black. Head length :
height : width ratio = 1 : 1.3 : 1.8. Scapus and pedicel uniformly yellow, without dark-
er marks, scapus with 5–6 rather long (as long as scapus) blackish setulae at dorso-api-
cal margin, pedicel with thin and short black setulae: 5–6 on dorso-lateral, 18–20 on
dorso-medial surface (1 longer seta plus 6–7 irregular rows of shorter setulae) and some
setulae forming single row at apico-ventral margin (not visible as antenna closely ap-
pressed to face). Flagellomere 1 light yellow, 1.5 times longer than wide. Arista bare,
8 V. A. Korneyev
Fig. 3. Orellia tragopogonis (1–6) and O. scorzonerae (7–8). 1 — head, lateral view; 2 — wing; 3 — fore femur
and tibia; 4, 7 — aculeus; 5, 8 — apex, enlarged; 6 — spermatheca (1 of the 2). Scale bar 1 mm.
Ðèñ. 3. Orellia tragopogonis (1–6) è O. scorzonerae (7–8). 1 — ãîëîâà, âèä ñáîêó; 2 — êðûëî; 3 — ïåðåäíåå
áåäðî è ãîëåíü; 4, 7 — àêóëåóñ; 5, 8 — âåðøèíà, óâåëè÷åíî; 6 — ñïåðìàòåêà (1 èç 2). Ìàñøòàáíàÿ
ëèíåéêà 1 ìì.
2.1 times longer than flagellomere 1, yellow in basal 0.25 of its length, dark brown in
the rest. Distance between bases of antennae 0.5 of scapus width. Palpus not extending
beyond anterior oral margin, pale yellow, with white setulae in basal 0.8 and 4–5 black
setulae at apex. Proboscis fleshy, somewhat elondate, with rather fine yellow setulae.
Frons as long as wide at posterior margin, with almost parallel sides; frontal plates with
4–7 black setulae at posterior 0.1 of its length, and white setulose in remaining por-
tion, orbital plates with 6–8 white setulae. Parafacialium at narrowest place 0.6 as wide
as flagellomere 1. Facialium with 3–4 rows of short brown to black setulae and gena
with 2 rows of longer blackish setulae; 2–3 whitish and 1–3 black setulae on genal ridge
in front of and above genal seta. Postgena with white setae below and behind of black
and long genal seta. Eye vertical, 1.5 times higher than long. Gena narrow, 0.14 as long
as eye vertical diameter. Head setae dark brown to black, except for postocellar, post-
vertical, postocular and occipital setae all white.
Thorax densely white setulose, with setae dark brown to black. Mesonotum dull,
short, but densely microtrichose, with black lyrate pattern and. Medial vitta reaching at
most dorsocentral setae posteriorly. Three pairs of shining black spots: at suture and at
bases of dorsocentral and acrostichal setae. Scutellum yellow, flattened. with 12–15
white setulae at each side and 4 setae; no brown or black spots around sockets of setae;
antero-ventral corners at squamal ridge blackish. Pleura subshining yellow; katepister-
num with large triangular dull black spot; meron at middle, katatergite and anatergite
at posteroventral margin dull black. Postnotum dull black, shining medially.
Wing (fig. 3, 2) with 4 yellow, brownish-bordered, partially joined crossbands. Vein
R4+5 with 1–3 setulae basally on dorsal surface and 0–1 on ventral surface. Basicostal,
costal and basal cubital cells completely, and basal radial cell to the level of BM-CU
vein level, brownish-yellow. Subbasal crossband restricted reaching A1 vein. Basal
medial cell with translucent window without microtrichiae. Subcostal cell brownish-yel-
low; subbasal and discal crossbands joined to R4+5 level; preapical crossband joined to
apical in cell r1 and partially in r2+3 cell. Postero-apical lobe of bcu reaching or very
slightly exceeding BM-Cu vein level.
Legs yellow, fore femur with 30–35 white setae arranged in 2–3 rows on dorsal
surface and one row of strong black setae on postero-ventral surface (fig. 3, 3).
Abdominal tergites all mixed black and white setulose, except tergite 1 white setu-
lose and tergites 5 and 6 with a few white setulae antero-medially. Tergites 2–5 with 4
black spots at anterior margin each. Tergites 3–5 white setulose with single row of black
marginal setae, tergite 6 with 7–8 long marginal setae, 1.8 times as long as tergite 5.
Tergosternite 7 yellow with 2 inconspicuous brownish spots basally, black setulose,
slightly longer than tergites 3–6 together. Aculeus bluntly tapered apically (fig. 3, 4, 5),
twice longer than costal cell.
Measurements (length in mm). Body (with oviscape) 7.0–7.8, wing 5.1–5.7, ovis-
cape dorsally 1.7–2.0, aculeus 2.2–2.44.
Ma l e not known.
B i o l o g y. Larvae feed in flower heads of Tragopogon dubium and pupate in a co-
coon made of remainders of pappi.
Commen t s. This species was first recognized as a new one by Dr. Jan Dirlbek
(Prague), who supplied me with the material from his personal collection and turned
my attention to some of its diagnostical characters.
The new species can be distinguished from other species of the genus Orellia by
the following key.
9New and Little-Known Tephritidae from Europe
Key to species of genus Orellia
Òàáëèöà äëÿ îïðåäåëåíèÿ âèäîâ ðîäà Orellia
1. Place of insertion of presutural supraalar and scutellar setae with brown or black spots ............ 2.
— Place of insertion of presutural supraalar and scutellar setae uniformly yellow, without brown or
black spots .................................................................................................................................... 3.
2. Discal and preapical crossbands separated. Katepisternum with black spot. Larvae in stems of
Tragopogon spp. (Europe and western Asia) .................................................... O. falcata (Scopoli)
— Discal and preapical crossbands joined. Katepisternum with reddish, rarely black, spot. Larvae in
flower heads of Tragopogon spp. or, rarely, in Scorzonera spp. and Taraxacum serotinum (Europe
and some regions of western and eastern Asia) ........... O. stictica (Gmelin) (= punctata Schrank)
3. Discal and preapical crossbands joined (fig. 3, 2). Fore femur dorsally with white setulae and setae,
except for very apex (fig. 3, 3). Oviscape slightly shorter than all abdominal tergites together.
Aculeus 2.2–2.44 mm long, 2 times as long as cell c (fig. 3, 4, 5). Larvae in flower heads of
Tragopogon (Spain) .................................................... O. tragopogonis Korneyev et Dirlbek, sp. n.
— Preapical and apical crossbands separated in 95% of specimens, and if joined, then fore femur
dorsally with black setulae and setae, (except for very base). Oviscape slightly longer than
abdominal tergites 4–6 together. Aculeus 1.4–1.6 mm long, 1.7–1.85 times as long as cell c (fig. 3,
7, 8). Larvae in flower heads of Scorzonera spp. (Europe, on wet meadows). ................................
................................................................... O. scorzonerae (Robineau-Desvoidy) (= distans Loew)
Chaetorellia australis Hering, 1937
Ma t e r i a l e x am in ed. Ukraine, 46°56'N, 31°25'E, near Grigorivka, swept from Centaurea cyanus L.,
20.07.1997, 2 {, 2 } (V. & S. Korneyev) (CVK).
Commen t s. First reliable record from Ukraine.
Chaetostomella rossica Hendel, 1927
Ma t e r i a l e x am in ed. SW of European Russia, Voronezh Oblast', Khoper Natural Reserve,
27.06.1987, 27.07.1987, {, 4 } (Kasparyan) (ZISP).
Commen t s. This species was primarily known only from its type locality (Sarep-
ta) and then recorded from Ukraine (Korneyev, 1985; 1987).
Urophora pauperata (Zaitzev, 1945)
Ma t e r i a l e x am in ed. Albania, Kula Ljums, 7–14.06.1918, } (Alban. Exp.) (NHMW).
Commen t s. First record from Albania.
Tephritis frauenfeldi Hendel, 1927
Ma t e r i a l e x am in ed. Greece, Olympus near Litohoro, h=800 m, ex flower heads of Jurinea mollis,
8.06.2002 — exit 18.06.2002, 30 {, 35 } (V. & S. Korneyev) (CVK; SIZK).
Commen t s. First record from Greece.
Tephritis hurvitzi Freidberg, 1982
Ma t e r i a l e x am in ed. Ukraine, Crimea, vicinity of Rybachye, Kanaka Valley, 29.04.1992; },
7–20.05.1993, 2 { (Korneyev) (CVK; SIZK).
Commen t s. First record from Ukraine.
Tephritis divisa Rondani, 1870
Ma t e r i a l e x am in ed. Ukraine, Crimea, vicinity of Rybachye, Kanaka Valley, 4.05.1992, 7 },
7–20.05.1993, { (Korneyev) (CVK; SIZK).
Commen t s. First record from Ukraine.
10 V. A. Korneyev
Tephritis arnicae (Linnaeus, 1758)
Hering, 1942; Richter, 1970: 155; Foote, 1984: (Orellia).
Ma t e r i a l e x am in ed. Moldova, “6 km NW Kotovsk” [= Hincesti], Loganesti forest, ex Doronicum
flower heads, 16.06 — exit 24.06.1987, 4 }, (Korneyev) (CVK).
Commen t s. First record from Moldova.
Xyphosia miliaria Hendel, 1927
Ma t e r i a l e x am in ed. Greece, Epirus, 24 km NNW of Ioannina, Monodendri, 39°52,9'N,
20°45,132'E, Vikos Canyon, 14.06.2002, 2 {, } (Korneyev) (CVK).
Commen t s. First record from Greece.
Myopites apicatus Freidberg, 1980
Ma t e r i a l e x am in ed. Bulgaria, Aladza monastyr, 08.1959, 4 } (J. Dirlbek); Kiten, oak wood + pig
farm, 42°14' N, 27°52' E, 17.07.1987, 3 {, }, Georgyi Trajkov, edge of oak wood, 43°00' N, 27°48' E, 07.1987,
{, Achtopol, pasture, 42°09', 27°52', 19.07.1987, 2 {, }, Sliven, along brook, 42°45', 26°17', 21.07.1987, 2{
(Bartak) (NMP); Ukraine: Crimea: Alupka, 23.09.1936, } (Filippov) (ZMUM).
Commen t s. First record from Bulgaria and Ukraine.
Myopites stylatus (Fabricius, 1794)
Ma t e r i a l e x am in ed. Albania: Golem-Kavaja, 09.1960, { (Dirlbek) (NMP).
Commen t s. First record from Albania.
Noeeta bisetosa Merz, 1992
Ma t e r i a l e x am in ed. Kyiv, Khodosievka, ex Hieracium sp., 6.08 — exit 15.09.1985, } (Korneyev)
(SIZK).
Commen t s. This species was originally described from Switzerland and then fo-
und in European Russia and Hungary (Basov, 1999; Merz, 2000)
Noeeta crepidis Hering, 1936
Commen t s. Re-examination of the material has shown that the record from
Moscow Region (Korneyev, 1982) was based upon a misidentified specimen of N. pu-
pillata (Fallén).
Xanthomyia alpestris (Pokorny, 1887)
Foote, 1984 (Paracarphrotricha); Norrbom et al., 2000 (Xanthomyia). — pseudoradiata Becker: Rich-
ter, 1970; Foote, 1984 (Paracarphrotricha).
Ma t e r i a l e x am in ed. Latvia, Berzs, "plava" [meadow], 18.06.1991, { [Karps leg.] (ZISP).
Commen t s. First record from Latvia.
Oxyna flavipennis (Loew, 1844)
Ma t e r i a l e x am in ed. Romania, Mehadia, 16.07.1912; {, Orsova, 6.06.1912, { (DEI).
Commen t s. First record from Romania.
I wish to express my sincere thanks to Bernhard Merz for reading this manuscript and useful critical comments.
Basov V. M. Tephritidae (Diptera) of Middle Volga and Cis-Ural areas [Áàñîâ Â. Ì. Ïåñòðîêðûëêè
(Diptera, Tephritidae) Ñðåäíåãî Ïîâîëæüÿ è Ïðåäóðàëüÿ // Âåñòí. Óäìóðòñ. óí-òà. Ñåð. Áèîëîã.
ðàçíîîáðàçèå Óäìóðòñêîé Ðåñïóáëèêè]. — 1999. — 2. — P. 33–45. — Russian.
Foote R. H. Family Tephritidae (Trypetidae) // Á . Soós, L. Papp (eds). Catalogue of Palaearctic Diptera.
Vol. 9. Micropezidae — Agromyzidae. — Budapest ; Amsterdam. — 1984. — P. 66–149.
11New and Little-Known Tephritidae from Europe
Korneyev V. A. On the fruit fly fauna (Diptera, Tephritidae) of the European Territory of the USSR [Êîðíå-
åâ Â. À. Ê ôàóíå ìóõ-ïåñòðîêðûëîê (Diptera, Tephritidae) åâðîïåéñêîé ÷àñòè ÑÑÑÐ] // Âåñòí.
çîîëîãèè. — 1982. — ¹ 2. — Ñ. 83–84. — Russian.
Korneyev V. A. Fruit flies of the tribe Terelliini Hendel, 1927 (Diptera: Tephritidae) of the fauna of the
USSR. [Êîðíååâ Â. À. Ìóõè-ïåñòðîêðûëêè òðèáû Terelliini Hendel, 1927 (Diptera, Tephritidae)
ôàóíû ÑÑÑÐ] // Ýíòîìîë. îáîçð. — 1985. — 64, âûï. 3. — P. 626–644. — Russian.
Korneyev V. A. Little known species of Tephritidae (Diptera) of the Ukrainian fauna [Êîðíååâ Â. À.
Ìàëîèçâåñòíûå âèäû ìóõ-ïåñòðîêðûëîê (Diptera, Tephritidae) ôàóíû Óêðàèíû] // Ôàóíà è
áèîöåíîòè÷åñêèå ñâÿçè íàñåêîìûõ Óêðàèíû. — Êèåâ : Íàóê. äóìêà. — 1987. — P. 83–87. —
Russian.
McAlpine J. F. Morphology and terminology // McAlpine J. F., Peterson B. V., Shewell G. et al.(coords.).
Manual of Nearctic Diptera. Vol. 1. — Ottawa : Research Branch, Agriculture Canada. — 1981. —
P. 9–63. — (Monograph of the Biosystematics Research Institute, No. 27)
Merz B. Additions and corrections to the checklist of Tephritidae of Hungary (Diptera: Acalyptrata) // Folia
Ent. Hung. — 2000. — 61. — P. 207–214.
Richter V. A. Family Tephritidae (Trypetidae) — fruit flies [Ðèõòåð Â. À. Tephritidae (Trypetidae) —
ïåñòðîêðûëêè // G. Y. Bei-Bienko, ed., Îïðåäåëèòåëü íàñåêîìûõ åâðîïåéñêîé ÷àñòè ÑÑÑÐ.
Òîì5. Äâóêðûëûå, áëîõè. ×. 2 [Keys to the insects of the European part of the USSR. Vol. 5.
Diptera, Siphonaptera. Part 2]. (Îïðåäåëèòåëè ïî ôàóíå ÑÑÑÐ, èçäàâàåìûå ÇÈÍ ÀÍ ÑÑÑÐ [Keys
to the USSR fauna published by ZIN AN SSSR]. No. 103). — Leningrad: Nauka. — 1970. —
P. 132–172. — Russian.
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