Saussurea porcii Degen (Asteraceae) in the Svydovets mountains (Ukrainian Carpathians): location, coenotic conditions, population parameters and conservation
Повідомлено про знахідку Saussurea porcii на Свидовці в Українських Карпатах. Вид трапляється у двох близько розташованих вологих кальценосних осередках у долині струмка по-між горами Трояска і Татарука на висоті близько 1300 м над р.м. Вказане точне місцезна-ходження і фітоценотичні умови оселищ, а...
Збережено в:
| Дата: | 2007 |
|---|---|
| Автори: | , , |
| Формат: | Стаття |
| Мова: | English |
| Опубліковано: |
Інститут ботаніки ім. М. Г. Холодного НАН України
2007
|
| Теми: | |
| Онлайн доступ: | http://dspace.nbuv.gov.ua/handle/123456789/3785 |
| Теги: |
Додати тег
Немає тегів, Будьте першим, хто поставить тег для цього запису!
|
| Назва журналу: | Digital Library of Periodicals of National Academy of Sciences of Ukraine |
| Цитувати: | Saussurea porcii Degen (Asteraceae) in the Svydovets mountains (Ukrainian Carpathians): location, coenotic conditions, population parameters and conservation / Yu.Ya. Kobiv, M. Helesh, L. Borsukevich // Укр. ботан. журн. — 2007. — Т. 64, № 6. — С. 825-832. — Бібліогр.: 30 назв. — англ. |
Репозитарії
Digital Library of Periodicals of National Academy of Sciences of Ukraine| id |
irk-123456789-3785 |
|---|---|
| record_format |
dspace |
| spelling |
irk-123456789-37852009-07-13T12:00:26Z Saussurea porcii Degen (Asteraceae) in the Svydovets mountains (Ukrainian Carpathians): location, coenotic conditions, population parameters and conservation Kobiv, Yu.Ya. Helesh, M. Borsukevich, L. Геоботаніка, екологія, охорона рослинного світу Повідомлено про знахідку Saussurea porcii на Свидовці в Українських Карпатах. Вид трапляється у двох близько розташованих вологих кальценосних осередках у долині струмка по-між горами Трояска і Татарука на висоті близько 1300 м над р.м. Вказане точне місцезна-ходження і фітоценотичні умови оселищ, а також основні популяційні та індивідуальні параметри. Популяція є нечисленною, знаходиться під загрозою і потребує охорони. Наведені результати поглиблюють відомості про поширення та екологічні потреби S. porcii — вкрай рідкісного реліктового виду, ендемічного для Східних Карпат. Сообщается о находке Saussurea porcii на Свидовце в Украинских Карпатах. Вид встречается в двух близлежащих влажных кальценосных местообитаниях в долине ручья между горами Трояска и Тататрука на высоте около 1300 м над у.м. Указывается точное расположение и ценотические условия местопроизрастаний, а также основные популяционные и индивидуальные параметры. Популяция является малочисленной, находится под угрозой исчезновения и нуждается в охране. Приведенные результаты углубляют сведения о распространении и экологических потребностях S. porcii — крайне редкого реликтового вида, эндемического для Восточных Карпат. 2007 Article Saussurea porcii Degen (Asteraceae) in the Svydovets mountains (Ukrainian Carpathians): location, coenotic conditions, population parameters and conservation / Yu.Ya. Kobiv, M. Helesh, L. Borsukevich // Укр. ботан. журн. — 2007. — Т. 64, № 6. — С. 825-832. — Бібліогр.: 30 назв. — англ. 0372-4123 http://dspace.nbuv.gov.ua/handle/123456789/3785 en Інститут ботаніки ім. М. Г. Холодного НАН України |
| institution |
Digital Library of Periodicals of National Academy of Sciences of Ukraine |
| collection |
DSpace DC |
| language |
English |
| topic |
Геоботаніка, екологія, охорона рослинного світу Геоботаніка, екологія, охорона рослинного світу |
| spellingShingle |
Геоботаніка, екологія, охорона рослинного світу Геоботаніка, екологія, охорона рослинного світу Kobiv, Yu.Ya. Helesh, M. Borsukevich, L. Saussurea porcii Degen (Asteraceae) in the Svydovets mountains (Ukrainian Carpathians): location, coenotic conditions, population parameters and conservation |
| description |
Повідомлено про знахідку Saussurea porcii на Свидовці в Українських Карпатах. Вид трапляється у двох близько розташованих вологих кальценосних осередках у долині струмка по-між горами Трояска і Татарука на висоті близько 1300 м над р.м. Вказане точне місцезна-ходження і фітоценотичні умови оселищ, а також основні популяційні та індивідуальні параметри. Популяція є нечисленною, знаходиться під загрозою і потребує охорони. Наведені результати поглиблюють відомості про поширення та екологічні потреби S. porcii — вкрай рідкісного реліктового виду, ендемічного для Східних Карпат. |
| format |
Article |
| author |
Kobiv, Yu.Ya. Helesh, M. Borsukevich, L. |
| author_facet |
Kobiv, Yu.Ya. Helesh, M. Borsukevich, L. |
| author_sort |
Kobiv, Yu.Ya. |
| title |
Saussurea porcii Degen (Asteraceae) in the Svydovets mountains (Ukrainian Carpathians): location, coenotic conditions, population parameters and conservation |
| title_short |
Saussurea porcii Degen (Asteraceae) in the Svydovets mountains (Ukrainian Carpathians): location, coenotic conditions, population parameters and conservation |
| title_full |
Saussurea porcii Degen (Asteraceae) in the Svydovets mountains (Ukrainian Carpathians): location, coenotic conditions, population parameters and conservation |
| title_fullStr |
Saussurea porcii Degen (Asteraceae) in the Svydovets mountains (Ukrainian Carpathians): location, coenotic conditions, population parameters and conservation |
| title_full_unstemmed |
Saussurea porcii Degen (Asteraceae) in the Svydovets mountains (Ukrainian Carpathians): location, coenotic conditions, population parameters and conservation |
| title_sort |
saussurea porcii degen (asteraceae) in the svydovets mountains (ukrainian carpathians): location, coenotic conditions, population parameters and conservation |
| publisher |
Інститут ботаніки ім. М. Г. Холодного НАН України |
| publishDate |
2007 |
| topic_facet |
Геоботаніка, екологія, охорона рослинного світу |
| url |
http://dspace.nbuv.gov.ua/handle/123456789/3785 |
| citation_txt |
Saussurea porcii Degen (Asteraceae) in the Svydovets mountains (Ukrainian Carpathians): location, coenotic conditions, population parameters and conservation / Yu.Ya. Kobiv, M. Helesh, L. Borsukevich // Укр. ботан. журн. — 2007. — Т. 64, № 6. — С. 825-832. — Бібліогр.: 30 назв. — англ. |
| work_keys_str_mv |
AT kobivyuya saussureaporciidegenasteraceaeinthesvydovetsmountainsukrainiancarpathianslocationcoenoticconditionspopulationparametersandconservation AT heleshm saussureaporciidegenasteraceaeinthesvydovetsmountainsukrainiancarpathianslocationcoenoticconditionspopulationparametersandconservation AT borsukevichl saussureaporciidegenasteraceaeinthesvydovetsmountainsukrainiancarpathianslocationcoenoticconditionspopulationparametersandconservation |
| first_indexed |
2025-07-02T07:01:43Z |
| last_indexed |
2025-07-02T07:01:43Z |
| _version_ |
1836517638838157312 |
| fulltext |
ISSN 0372�4123. Укр. ботан. журн., 2007, т. 64, № 6 825
Yu.Ya. KOBIV 1, M. HELESH 2, L. BORSUKEVICH 2
1 Institute of Ecology of the Carpathians,
National Academy of Sciences of Ukraine
Kozelnytska St., 4, Lviv, 79026, Ukraine
rehablv@link.lviv.ua
2 Botanical Garden of the I. Franko Lviv National University,
Cheremshyna St., 44, Lviv, 79014, Ukraine
botsad@franko.lviv.ua
SAUSSUREA PORCII DEGEN (ASTERACEAE)
IN THE SVYDOVETS MOUNTAINS
(UKRAINIAN CARPATHIANS): LOCATION,
COENOTIC CONDITIONS, POPULATION
PARAMETERS AND CONSERVATION
K e y w o r d s: Saussurea porcii, Eastern Carpathians, endemic, threatened
species, population, conservation
Abstract
A new find of Saussurea porcii is reported from the Svydovets Mountains, Ukrainian
Carpathians. It occurs on two adjacent wet calciferous sites in the stream valley between
Troyaska and Tataruka Mts. at the altitude about 1300 m a.s.l. Exact location and coeno�
tic conditions of the localities are described, as well as main population and individual
parameters. The population is low�numbered, endangered and needs conservation. Pre�
sented results extend the knowledge on distribution and ecological needs of S. porcii — an
extremely rare relict species endemic to the Eastern Carpathians.
Introduction
The Carpathians are considered one of the centers of biodiversity and endemism on Pan�
European scale [10]. Ukraine shares a significant part of the Eastern Carpathians, but the
regions of the highest biodiversity are situated mainly on the territory of Romania where
endemic taxa are most abundant [6, 14, 22]. This is caused by more severe Pleistocene
cooling that took place the Ukrainian Carpathians and their rather uniform geological
structure, which is presented mostly by sandstone flysch. Therefore, the occurrence of
scarce local endemics in the Ukrainian Carpathians is of special interest.
Perhaps, the most remarkable of such narrow East�Carpathian endemics is Saussu�
rea porcii Degen, which was even called «half�legendary» [28: 124] and «the rarest of all
known Carpathian plants» [8: 141]. It was discovered in 1856 by F. Porcius near Coron�
gisu Mt. in the Rodna massif, Transylvanian part of the Eastern Carpathians on the terri�
tory of today's Romania (fig. 1, locality 1). Initially the plant was determined as Saussurea
serrata DC. and that caused a sensation among the botanists, because the latter species
was known mostly from Siberia. Later on A. Degen after detailed studies made a conclu�
sion that the Carpathian plants present a separate species, which he called Saussurea
porcii [12]. However, close affinity of S. porcii with a Siberian species S. parviflora (Poir.)
DC. (= S. serrata) is still accepted [3, 4].
© Yu.Ya. KOBIV, M. HELESH, L. BORSUKEVICH, 2007
ISSN 0372�4123. Ukr. Botan. Journ., 2007, vol. 64, № 6826
In the 1920—1930 Saussurea porcii was also found in the Chyvchyny [29, 30] and
Chornohora Mts. [15], i.e. on the territory of the Ukrainian Carpathians (fig. 1, locali�
ties 2—4).
The main region of the species distribution lies in the Chyvchyny Mts. at the
headwaters of the White and Black Cheremosh rivers. The localities are restricted to
two places: 1) southernmost part of the Chyvchyny massif on the slopes of Hnetiesa
Mt. near the border with Romania and 2) about 11 km to the north on Hlystowaty Mt.
near the pass of Shyya between the Chyvchyny and Hryniava Mts. In both places
S. porcii occurs in several localities that suggests the existence of two metapopulations
in the region. The localities are confined to peat bogs situated within 1380—1560 m
a.s.l. [24].
In the Chornohora Mts. the species is known from the south�western foothill of
Petros Mt. where it occurs on a mown fen meadow about 1350—1400 m a.s.l. [1, 2].
Recently we managed to find a new locality of the species in the Svydovets Mts. and
further material presents the obtained data (fig. 1, lpcality 5; fig. 2).
Saussurea porcii is included into «European Red List …» [16], «Red Data Book of
Ukraine» [7], Romanian «Red List» [13] and is a species of top interest from the biogeo�
graphical and conservation points of view. It is considered extinct in the only known
Romanian locality, i.e. in locus classicus in the Rodna Mts. [11, 13, 27].
Fig. 1. Distribution of Saussurea porcii Degen. S y m b o l s i n d i c a t e : —
mountains; — localities (1 — in the Rodna (extinct), 2—3 — Chyvchyny, 4 —
Chornohora, 5 — Svydovets Mts. — newly found)
Material and Methods
The results presented below were obtained during field research performed in the central
part of the Svydovets massif in July, 2005. Using meander search [17] we looked for loca�
lities of rare and threatened plant species and paid special attention to calciferous habitats.
The exact location of the site was determined in WGS�84 system with Garmin eTrex
Global Positioning System (GPS) navigator with the accuracy of measurements within
10 metres.
A few individuals of Saussurea porcii were transplanted to the Botanical Garden of
the I. Franko Lviv National University to provide off�site conservation of genetic pool of
the population within the collection of critically endangered species in case of extinction
in situ.
Species abundance was evaluated according to the grades of J. Braun�Blanquet's
scale [9].
Much of the above information on the distribution of Saussurea porcii in the
Chyvchyny Mts. was obtained from personal collections of B. Pawlowski and J. Ma�
dalski kept in the Herbarium of the W. Szafer Institute of Botany (KRAM) in Krakow,
Poland.
ISSN 0372�4123. Укр. ботан. журн., 2007, т. 64, № 6 827
Fig. 2. Location of S. porcii in the Svydovets Mts. S y m b o l s i n d i c a t e: —
species localities; I, II — numbers of the described stations
Collected specimens are deposited in the Herbarium of the M.G. Kholodny Institu�
te of Botany (KW) in Kyiv and the Herbarium of the I. Franko Lviv National Universi�
ty (LW).
Results
Location of the site and its coenotic conditions. A population of Saussurea porcii was discov�
ered in the place of Podyna on two neighbouring stations in the stream valley between
mountains of Troyaska and Tataruka in the central part of the Svydovets massif just on the
border between Rakhiv and Tiachiv Districts, Transcarpathian (Zakarpattya) Region (fig. 2).
Station I is restricted to the peat bog lying on the gentle slope at the bottom of the
north�eastern foothill of Troyaska Mt. at the altitude of 1295 m a.s.l. The slope has a
northern aspect with a mean angle of about 3°. Geographical coordinates of the locality
are: 48°17′30.5′′N; 24°09′23.4′′E.
The area of the site is about 400 m2. Herb layer covers about 60 % of it, while moss�
es account for another 30 %. The species composition of vascular plants with correspon�
ding grades of abundance is as follows: Saussurea porcii — +, Carex paniculata L. — 3,
C. hartmannii Cajander — 3, C. flava L. — 1, C. panicea L. — +, C. echinata Murray — +,
Swertia perennis L. subsp. alpestris (Baumg. ex Fuss) Simonk. — 1, Eriophorum latifolium
Hoppe — 1, Molinia caerulea (L.) Moench — 1, Astrantia major L. — +, Dactylorhiza
cordigera (Fries) Soo — +, Equisetum sylvaticum L. — +, Alnus viridis (Chaix) DC. — +,
Pinguicula vulgaris L. — +, Potentilla erecta (L.) Rauchel — +, Prunella vulgaris L. — +,
Salix silesiaca Willd. — +, Trollius altissimus Crantz — +; and bryophytes: Campylium
stellatum (Hedw.) J. Lange & C. Jens. — 3, Drepanocladus aduncus (Hedw.) Warnst. — 2,
Rhytidiadelphus triquetrus (Hedw.) Warnst. — 2, Fissidens cristatus Wils. — 1.
The soil is neutral with the pH (H2O) value about 6.5. Present conditions of high
moistening are most favourable for hygrophytous vegetation.
Station II is situated about 150 m to the west from the previous one at the altitude of
1300 m a.s.l. on the left bank of the stream that flows down from the cirque between
Troyaska and Tataruka Mts. The aspect of the slope is south�eastern and its inclination is
about 5°. The station has such geographical coordinates: 48°17′31.1′′N; 24°09′18.1′′E.
The area of the site is only 25 m2. Herbaceous and shrub vegetation covers about 90 %.
Following species are presented here: Saussurea porcii — +, Carex paniculata — 4, C. syl�
vatica Hudson — 1, C. sempervirens Vill. — 1, C. pallescens L. — 1, C. panicea — +, Mo�
linia caerulea — 2, Equisetum sylvaticum — 1, Equisetum palustre L. — +, Filipendula ul�
maria (L.) Maxim. — 1, Salix silesiaca — 1, Astrantia major — +, Alnus viridis — +, Cam�
panula serrata (Kit.) Hendrych — +, Crepis paludosa (L.) Moench — +, Cruciata glabra
(L.) Ehrend. — +, Dactylorhiza cordigera — +, Potentilla erecta — +, Prunella vulgaris —
+, Swertia perennis subsp. alpestris — +, Trollius altissimus — +; Valeriana simplicifolia
Kabath — +, Veratrum album L. — +, Viola biflora L. — +. In general, herbaceous vege�
tation is secondary and appeared in place of former Salici�Alnetum viridis schrub com�
munity, which had stretched along the stream, but was later destroyed by human activity
that accompanied long�term grazing in the region. The patches of Alnus viridis and Salix
silesiaca still remain in close vicinity with the site.
ISSN 0372�4123. Ukr. Botan. Journ., 2007, vol. 64, № 6828
The pH (H2O) value of the soil is 6.6, i.e. almost the same as in the previous site.
This station, however, is much drier and its conditions may be characterized as hygrome�
sophytous.
Both sites belong to Caricetum paniculatae association (Magnocaricion alliance). In
addition to S. porcii, some other rare and threatened calciphilous taxa occur here as well,
e.g. Swertia perennis subsp. alpestris, Carex hartmannii and Pinguicula vulgaris. However,
the species composition in the Svydovets locality is poorer than in the habitats of Saus�
surea porcii known from the Chyvchyny Mts. where Festuca porcii Hackel and F. apenni�
na De Not occur that are characteristic for Cariceto�Festucetum porcii association (Carex
paniculata & Festuca porcii), which is endemic to the Eastern Carpathians [23, 24].
Analysis of the data from literature shows that Carex paniculata occurs in all its known
from the Ukrainian Carpathians localities and dominates in most of them. Other species
common to all the localities are: Carex flava, C. echinata, Dactylorhiza cordigera, Filipen�
dula ulmaria, Potentilla erecta and Astrantia major [1, 2, 24].
Population and individual parameters. Both described above sites present a single
metapopulation of Saussurea porcii, which consists of two smaller subpopulations inha�
biting stations I and II.
Four fertile and 7 sterile individuals were found in station I, while the number of
these groups in station II was 12 and 20 respectively. Thus, the overall population number
is very low and makes 43 individuals. This may shaw that the population is endangered.
Rather low percentage of sterile individuals within the whole population (62.8 %) may be
caused by poor recruitment via seed reproduction.
Due to clonal growth and rhizome branching each mature individual consists of sev�
eral modules. A fertile individual has up to 5 flower stems and 1—3 sterile shoots. Rhizo�
matous growth is sympodial and annual increments make 10 to18 mm. On the late stage
of the life history clonal disintegration occurs. However, it is accompanied by senescence
and does not play an important role in sustainment of the population. Height of fertile
shoots ranges within 45—74 cm (58.5±2,5 cm on average) and of sterile ones — within
18—28 cm. The number of capitula per flower stem is 8 to 34. The vitality of individuals
is significantly lower than in plants from Hlystovatyi in the Chyvczyny Mts. where
according to our data from 2004 the height of fertile individuals ranged within 53—98 cm.
The Svydovets population is the least numerous in comparison with the other species
localities from the Ukrainian Carpathians, i.e. from the Chyvchyny and Chornohora Mts.
as appears from the published data [1, 2, 24] and our former field observations. Popula�
tion viability is low and reliable prognosis on its survival in the long�term perspective is
problematic. The above data lay down a landmark for future monitoring the population.
Most probably, the stunted condition of the population is caused by severe anthro�
pogenic impact, namely grazing and accompanying trampling that took place in the past
and had a significant influence upon the vegetation all around. However, grazing has
almost stopped last years and will hardly restore in future because cattle breeding became
unprofitable in the remote mountain regions of the Ukrainian Carpathians like all over
Europe. Possibly, the Svydovets population is recovering now and will gradually increase,
especially on its indigenous peat bog habitat in station I.
ISSN 0372�4123. Укр. ботан. журн., 2007, т. 64, № 6 829
On the other hand, the relict character of the species, its extreme rarity and extinc�
tion in the Romanian locality may suggest that it is prone to natural decline [11].
Conservation considerations. Saussurea porcii is a very stenotopic species and a set of
quite special habitat conditions is needed to provide viability of its populations. According
to our observations and published data [1, 23], all known populations of the species are
restricted to calciferous peat bogs or moist meadows within 1295�1560 m a.s. l. The habi�
tats that meet all these demands (in high content of calcium in soil, proper moistening
and narrow altitudinal range) are extremely rare in the Ukrainian Carpathians. They may
be treated as so�called «habitat islands», i.e. the sites with very special conditions differ�
ing from their surrounding [19, 20, 26]. Probability of a colonization event of such an
«island» from similar rather remote sites is very low. This explains the fact that S. porcii
does not occur in all the suitable habitats.
Localities of S. porcii are of great conservation value, because they harbour many
other species included into the «Red Data Book of Ukraine» [7] occur there, e.g. Festuca
porcii, Carex buxbaumii, Carex davalliana, Dactylorhiza cordigera, Pinguicula vulgaris and
Swertia perennis subsp. alpestris [1, 2, 24]. The three latter taxa occur in the described
above Svydovets locality, as well as a rather rare calciphilous sedge Carex hartmannii.
Thus, the discovered locality deserves conservation that could be provided by includ�
ing it into the Svydovets massif of the Carpathian Biosphere Reserve. In addition to the
place of Podyna, where S. porcii occurs, the whole stream valley and the cirque between
Troyaska and Tataruka Mts. deserve the status of the core zone, because that territory is
remarkable for its biodiversity. Very numerous populations of Gentiana lutea L., Narcissus
angustifolius Curtis, Pinguicula vulgaris and Swertia perennis subsp. alpestris — taxa from
the «Red Data Book of Ukraine» [7] — occur here.
Conclusion
The discovery of Saussurea porcii in the Svydovets Mts. allows to revise some of the exist�
ing opinions concerning that species. First, its localities are not that unique and the
species range is somewhat wider than it was considered and includes four massifs of the
Eastern Carpathians — the Rodna, Chyvchyny, Chornohora and Svydovets Mts. The
main part of the range and all its known remaining localities are situated on the territory
of Ukraine (fig. 1). The largest distance between localities, i.e. in the Rodna Mts. (where
the species is already extinct) and in the Svydovets Mts. is about 110 km.
Second, referring S. porcii to high�mountain or subalpine species [5, 20, 28] is not
quite appropriate that especially concerns the discovered site from the Svydovets Mts. sit�
uated at 1295—1300 m a. s. l. Actually all the species localities are restricted to the altitu�
dinal zone of mountain spruce forests, which in the biogeographical aspect correspond to
boreal forests of Siberia, where lies the main part of the range of a closely related species
Saussurea parviflora. The latter species occurs mostly in Western and Eastern Siberia,
Northern Mongolia and Nort�Eastern China, but its westernmost localities are also scat�
tered in European part of Russia about 300 km to the west from the Urals [3, 4, 22, 23].
The distance between these stations and the localities of S. porcii in the Carpathians is
very large and makes about 2400 km. Such a considerable disjunction in the range of these
ISSN 0372�4123. Ukr. Botan. Journ., 2007, vol. 64, № 6830
ISSN 0372�4123. Укр. ботан. журн., 2007, т. 64, № 6 831
two species as well as there morphological differentiation suggest a rather early time of
their divergence. Most authors refer S. porcii to Tertiary relicts [8, 24, 26, 29]. However,
taking into account rather oligothermic conditions of its habitats and definite Siberian
genetic links B. Pawlowski suggested that it could have migrated to Europe during the
severest cooling in the Pleistocene [22, 61].
Possibly, the Carpathians are also a secondary center of diversity for other taxa of Si�
berian origin, e.g. Ligularia sibirica (L.) Cass. s. l. and Cortusa matthioli L. s. l. that de�
monstrate some variability in the region.
Acknow Cedgement. We are very grateful to Dr. Ihor Danylkiv from the Institute of
Ecology of the Carpathians, National Academy of Sciences of Ukraine for determining the
bryophytes from the studied locality.
1. Данилик І.М., Антосяк В.М. Carex davalliana Smith (Cyperaceae) — новий вид для флори
високогір'я Українських Карпат // Укр. ботан. журн. — 1997. — 54, № 3. — С. 275—277.
2. Ефремов А.П. О находке соссюреи Порциуса (Saussurea porcii Degen) в Украинских
Карпатах // Биол. науки. — 1987. — 11. — С. 101—102.
3. Липшиц С.Ю. Род 1582. Соссюрея, горькуша — Saussurea DC. // Флора СССР. — М.; Л.:
Изд�во АН СССР, 1962. — С. 361—537.
4. Липшиц С.Ю. Род Saussurea DC. — Л.: Наука, 1979. — 283 с.
5. Малиновський К., Царик Й., Кияк В., Нестерук Ю. Рідкісні, ендемічні, реліктові та
погранично�ареальні види рослин Українських Карпат. — Львів: Ліга�Прес, 2002. — 76 с.
6. Стойко С., Гадач Е., Шимон Т., Михалик С. Заповідні екосистеми Карпат. — Львів: Світ,
1991. — 247 c.
7. Червона книга України. Рослинний світ / За ред. Ю.Р. Шеляга�Сосонка. — К.: УЕ, 1996. —
С. 608.
8. Чопик В.I. Високогiрна флора Українських Карпат. — К.: Наук. думка, 1976. — 270 с.
9. Braun�Blanquet J. Pflanzensoziologie. — 2 Aufl. — Wien: Springer�Verlag, 1951. — 581 p.
10. Centres of plant diversity: A guide and strategy for their conservation / Eds. S.D. Davis, V.H. Hey�
wood, A.C. Hamilton. — Vol. 1. — Cambridge: WWF & IUCN, 1994. — 250 p.
11. Coldea G., Wagner I. Potential scenarios on mountain biodiversity in the Eastern Carpathians re�
lated to climate change // First Int. Conf. on Mountain Biodiversity (7—10 September, 2000):
Abstracts. — Rigi�Kaltbad, 2000. — P. 16—17.
12. Convention on conservation of European wildlife and habitats. Appendix 1 of May 1999: Strivtly
protected flora specias. — Bern, 1999. — 19 p.
13. Degen A. Bemerkungen uber einige orientalische Pflanzenarten. — Magyar Botanikai Lapok. —
1904. — III. — S. 311—320.
14. Dihoru G., Dihoru A. Plante rare, periclitate si endemice in flora Romaniei — Lista Ro ie //
Acta Bot. Horti Bucure tensis. — 1993—1994. — P. 173—197.
15. Dihoru G., P rvu C. Plante endemice n flora Romaniei. — Bucure ti: Ceres, 1987. — 181 p.
16. Domin K. Saussurea porcii Degen v Podkarpatsk Rusi // V da p irodni. — 1932. — 13. —
S. 216—217.
17. European Red List of globally threatened animals and plants. — New York: UN, 1991. — 154 p.
18. Given D.R. Principles and practice of plant conservation. — London; Glasgow; Weinheim:
Chapman & Hall, 1997. — 260 p.
19. Gorman M. Island ecology. — London: Chapman & Hall, 1979. — 90 p.
20. Kobiv Y. Calciferous headstream localities as sites of noteworthy biodiversity in the Chornohora
Mts (Eastern Carpathians) // Roczniki Bieszczadskie. — 2005. — 13. — P. 203—214.
21. Lipschitz S.I. Saussurea DC. // Flora Europaea. — Vol. 4. — Cambridge; New York; Melbour�
ne: Cambridge Univ. Press, 1976. — P. 216—217.
22. Meusel H., Ja
..
ger E.J. Vergleichende Chorologie der Zentraleuropa
..
ischen Flora. — B. III. —
Jena; Stuttgart; New York: G. Fischer Verlag, 1992. — 650 s.
23. Paw owski B. Og lna charakterystyka geobotaniczna g r Czywczynskich // Rozpr. wydz. mat.�
przyrod. PAU. — 1946. — 72. — Dz. B. — S. 1—75.
24. Paw owski B., Walas J. Les assotiations des plantes vasculaires des Monts de Czywczyn // Bull.
Acad. Pol. Scient. et Letter. — Ser. B. — V. 1. — P. 117—181.
25. Pax F. Grundzu
..
ge der Pflanzenverbreitung in den Karpathen. — Leipzig, 1898. — Vol. I. — 269 s.
26. Selvi F. Rare plants on Mount Amiata, Italy: vulnerability to extinction on an ecological
«island» // Biol. Conserv. — 1997. — 81. — P. 257—266.
27. So R. Analyse der Flora des historischen Ungarns (Elemente, Endemismen, Relikte) //
Magyar Biol. Kut. Int. — 1933. — VI. — S. 172—193.
28. Tasenkevich L. Vascular plants // Carpathian list of endangered species. —Vienna; Krakow:
Europress, 2003. — P. 8—19.
29. Wilczy ski T. Zapiski florystyczne z Karpat Pokuckich // Rozpr. i wiad. Muz. im. Dzieduszy�
ckich. — 1924. — 10. — S. 114—127.
30. Wilczy ski T. U
..
ber das Vorkommen der Saussurea porcii Deg. im Czeremosz�Quellegebiete //
Magyar Botanikai Lapok. — 1927. — XXVI. — S. 65—69.
Recommended for publication Submitted 24.01.2007
by S.L. Mosyakin
Ю. Кобів 1, М. Гелеш 2, Л. Борсукевич 2
1 Інститут екології Карпат НАН України, м. Львів
2 Ботанічний сад Львівського національного університету ім. І. Франка
SAUSSUREA PORCII DEGEN (ASTERACEAE)
НА СВИДОВЦІ (УКРАЇНСЬКІ КАРПАТИ): МІСЦЕЗНАХОДЖЕННЯ,
ЦЕНОТИЧНІ УМОВИ, ПОПУЛЯЦІЙНІ ПАРАМЕТРИ ТА ОХОРОНА
Повідомлено про знахідку Saussurea porcii на Свидовці в Українських Карпатах. Вид трап�
ляється у двох близько розташованих вологих кальценосних осередках у долині струмка по�
між горами Трояска і Татарука на висоті близько 1300 м над р.м. Вказане точне місцезна�
ходження і фітоценотичні умови оселищ, а також основні популяційні та індивідуальні пара�
метри. Популяція є нечисленною, знаходиться під загрозою і потребує охорони. Наведені
результати поглиблюють відомості про поширення та екологічні потреби S. porcii — вкрай
рідкісного реліктового виду, ендемічного для Східних Карпат.
К л ю ч о в і с л о в а: Saussurea porcii, Східні Карпати, ендемік, популяція, охорона.
Ю. Кобив 1, М. Гелеш 2, Л. Борсукевич 2
1 Институт экологии Карпат НАН Украины, г. Львов
2 Ботанический сад Львовского национального университета им. И. Франка
SAUSSUREA PORCII DEGEN (ASTERACEAE)
НА СВИДОВЦЕ (УКРАИНСКИЕ КАРПАТЫ): МЕСТОНАХОЖДЕНИЯ,
ЦЕНОТИЧЕСКИЕ УСЛОВИЯ, ПОПУЛЯЦИОННЫЕ ПАРАМЕТРЫ И ОХРАНА
Сообщается о находке Saussurea porcii на Свидовце в Украинских Карпатах. Вид встречает�
ся в двух близлежащих влажных кальценосных местообитаниях в долине ручья между гора�
ми Трояска и Тататрука на высоте около 1300 м над у.м. Указывается точное расположение и
ценотические условия местопроизрастаний, а также основные популяционные и индивиду�
альные параметры. Популяция является малочисленной, находится под угрозой исчезнове�
ния и нуждается в охране. Приведенные результаты углубляют сведения о распространении
и экологических потребностях S. porcii — крайне редкого реликтового вида, эндемическо�
го для Восточных Карпат.
К л ю ч е в ы е с л о в а: Saussurea porcii, Восточные Карпаты, эндемик, популяция, охрана.
ISSN 0372�4123. Ukr. Botan. Journ., 2007, vol. 64, № 6832
|